Merlin Tuttle

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Merlin Devere Tuttle, (born August 26, 1941) is an American ecologist, conservationist and wildlife photographer who has specialized in bat ecology, behavior and conservation. He was the founder of the conservation organizations, Bat Conservation International (BCI) [1] and Merlin Tuttle's Bat Conservation [2].

Tuttle is known for founding the conservation organization, Bat Conservation International (BCI), and his research on gray bat population ecology migration,[1][2][3] and frog-eating bats,.[4][5][6][7][8][9][10][11][12][13] He is also recognized for his photography of bats.[14][15]

Early life[edit]

External audio
Adventures With a Bat Biologist, 12:29, To the Best of Our Knowledge[16]

Tuttle entered graduate school at the University of Kansas. His master's degree thesis focused on zoogeography of Peruvian bats.[17] He obtained his Ph.D. in 1974, completing his thesis on population ecology and migration of gray bats. He subsequently published several academic papers based on his research.[18][19][20][21][22][23][24]


  1. ^ Tuttle, M.D. 1975. “Population ecology of the gray bat (Myotis grisescens): Factors influencing preflight growth and development.” Occ. Pap. Mus. Nat. Hist., Univ. Kans., 36:1-24.
  2. ^ Tuttle, M.D. 1976. “Population ecology of the gray bat (Myotis grisescens): Factors influencing growth and survival of newly volant young.” Ecology, 57:587-595.
  3. ^ Tuttle, M.D. 1976. “Population ecology of the gray bat (Myotis grisescens): Philopatry, timing and patterns of movement, weight loss during migration, and seasonal adaptive strategies.” Occ. Pap. Mus. Nat. Hist., Univ. Kans., 54:1-38.
  4. ^ Barclay, R.M., Fenton, M.B., Tuttle, M.D. and Ryan, M.J. 1981. “Echolocation calls produced by Trachops cirrhosis (Chiroptera: Phyllostomatidae) while hunting for frogs.” Can. J. Zool., 59:750-753.
  5. ^ Ryan, M.J., Tuttle, M.D. and Taft, L.K. 1981. “The costs and benefits of frog chorusing behavior.” Behav. Ecol. Sociobiol., 8:273-278.
  6. ^ Tuttle, M.D. and Ryan, M.J. 1981. “Bat predation and the evolution of frog vocalizations in the Neotropics.” Science, 214(4521):677-678.
  7. ^ Ryan, M.J. and Tuttle, M.D. 1982. “Bat predation and sexual advertisement in a Neotropical anuran.” Amer. Nat., 119:136-139.
  8. ^ Tuttle, M.D., Taft, L.K. and Ryan, M.J. 1982. “Evasive behavior of a frog in response to bat predation.” Anim. Behav., 30:393-397.
  9. ^ Tuttle, M.D. and Ryan, M.J. 1982. “The role of synchronized calling, ambient light, and noise in anti-bat-predator behavior of a tree frog.” Behav. Ecol. Sociobiol., 11:125-131.
  10. ^ Ryan, M.J., Tuttle, M.D. and Barclay, R.M.R. 1983. “Behavioral response of the frog-eating bat, Trachops cirrhosus to sonic frequencies.” J. Comp. Physiol., 150:413-418.
  11. ^ Ryan, M.J. and Tuttle, M.D. 1983. “The ability of the frog-eating bat to discriminate among novel and potentially poisonous species using acoustic cues.” Anim. Behav., 31:827-833.
  12. ^ Tuttle, M.D., Ryan, M.J. and Belwood, J.J. 1985. “Acoustical resource partitioning by two species of phyllostomatid bats (Trachops cirrhosus) and Tonatia sylvicola).” Anim. Behav., 33:1369-1270.
  13. ^ Ryan, M.J. and Tuttle, M.D. 1987. “The role of prey-generated sounds, vision and echolocation in prey localization by the African bat, Megaderma cor megadermatidae).” J. Comp. Physiol., 161:59-66.
  14. ^ Allen, W.L. 2001. National Geographic 100 Best Pictures (Collector’s Edition Vol.1). Nat. Geo. Soc. Pub.
  15. ^ 2007. “The best wildlife photographers of the world named by DP magazine in the year 2007.”
  16. ^ "Adventures With a Bat Biologist". To the Best of Our Knowledge via WNYC. August 28, 2016. Retrieved November 3, 2016. 
  17. ^ Tuttle, M.D. 1970. “Distribution and zoogeography of Peruvian bats with comments on natural history.” Univ. Kans. Sci. Bull., 49: 45-86.
  18. ^ Tuttle, M.D. and Robertson, P.B. 1969. “The gray bat, Myotis grisescens, east of the Appalachians.” J. Mammal., 50:37.
  19. ^ Tuttle, M.D. and Stevenson, D.E. 1977. “An analysis of movement as a mortality factor in the gray bat, based on public recoveries of banded bats.” Amer. Midland Nat., 97:235-240.
  20. ^ Tuttle, M.D. 1978. “Variation in the cave environment and its biological implications.” Pp. 108-121 in Natl. Cave Management Sympos. Proc., 1977 (R. Zuber, J. Chester, S. Gilbert and D. Rhodes, eds.). Adobe Press, Albuquerque, NM.
  21. ^ Tuttle, M.D. 1979. “Status, causes of decline, and management of the endangered gray bat. J. Wildl. Manag., 44:955-960.
  22. ^ Tuttle, M.D. 1982. “Gray Bat.” Pp. 127-128 in Handbook of Census Methods for Terrestrial Vertebrates. (D.E. Davis, ed). CRC Press, Boca Raton, Florida.
  23. ^ Humphrey, S.R. and Tuttle, M.D. 1978. “Myotis grisescens.” Pp. 1-3, in Rare and endangered biota of Florida. Fla. Aud. Soc. and Def. Envir., 1:1-52.
  24. ^ Stevenson, D.E. and Tuttle, M.D. 1981. “Survivorship in the endangered gray bat (Myotis grisescens).” J. Mammal., 62:244-257.

Additional Reading[edit]

  • Tuttle, Merlin. The Secret Lives of Bats: My Adventures With the World's Most Misunderstood Mammals. Boston: Houghton Mifflin Harcourt, 2015.

External links[edit]