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Temporal range: Middle JurassicLate Cretaceous 166–66 Ma
Specimen of Darwinopterus modularis
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Clade: Breviquartossa
Clade: Monofenestrata
et al., 2010

Monofenestrata is an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea.[1] The earliest known monofenestrate fossils have been found in the Stonesfield Slate formation (UK), which dates to the Bathonian stage of the Middle Jurassic, dated to about 166 million years ago.[2]


  1. ^ Lü, J., Unwin, D.M., Jin, X., Liu, Y. and Ji, Q. (2010). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull." Proceedings of the Royal Society B, 277(1680): 383-389. doi:10.1098/rspb.2009.1603 PMID 19828548
  2. ^ Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." Geological Magazine, (advance online publication) doi:10.1017/S0016756811001154