The Mydidae (alternative spelling Mydaidae), or Mydas flies, are a cosmopolitan family of flies. It is a small family, with about 471 species described. They are generally large in size, including, the largest known fly, Gauromydas heros (syn. Mydas heros). Many of the species, in addition to their large size, are mimics of stinging hymenopterans, especially wasps. Most mydids are found in arid and semiarid regions of the world, but they are also found in other habitats. They are infrequently encountered as the adult lifespan can be quite short. Little is known about their biology, though Zikan reported the larvae of Gauromydas heros live in the subterranean detritus "pans" of Atta ants in southern Brazil, where they appear to be feeding on detritivorous Dynastinae (Coelosis spp.) larvae. In the U.S., Mydas brunneus, Mydas clavatus, and Mydas tibialis larvae are predatory on deadwood-feeding scarab beetle larvae (Osmoderma spp.) and can be found in standing and downed trees with extensive heart rot. Others (e.g. Mydas maculiventris) are subterranean and feed on "white grubs" (Scarabaeidae: genus Phyllophaga) that attack the roots of grasses and could be potential biocontrol agents of white grubs in sod production areas. Larvae typically take two or three years to mature. Adults of several species are avid flower visitors and act as pollinating agents. Rattlesnake master (Eryngium yuccifolium) is a favorite nectar source in the Midwest.
For terms see Morphology of Diptera.
Mydids are medium-sized to very large flies (9–60 mm in body length). The abdomen is long and cylindrical in section. It is slightly tapered apically in the male, and usually widest at segment four in the female. The second segment of the antenna forms a club. Mydids are sparsely pilose, and lack bristles except on the legs. The hind leg is much longer and stronger than both the middle leg and the fore leg and the hind femur is usually swollen and bears ventral spines. The hind tibia has an apical spur or bristles. The wings are long, and narrow to wide. Most of the veins end in the upper margin before the apex.
The classification of the family has changed fairly recently, with the inclusion of a few genera that were previously placed in the family Apioceridae. Apiocerids have long been given the common name "flower-loving flies", but only the group of "flower-loving flies" that actually visit flowers were transferred to the Mydidae. Among the genera transferred is Rhaphiomidas, which includes one of the few Diptera on the United States' Endangered Species List, the Delhi Sands flower-loving fly (Rhaphiomidas terminatus abdominalis).
- Lyons, Kathleen; Dikow, Torsten (29 December 2010). "Taxonomic revision of Ectyphus Gerstaecker, 1868 and Parectyphus Hesse, 1972 with a key to world Ectyphinae (Insecta, Diptera, Mydidae)". ZooKeys. 73: 25–59. PMC . doi:10.3897/zookeys.73.840.
- Calhau, Julia; Lamas, Carlos José Einicker; Nihei, Silvio Shigueo (2015). "Review of the Gauromydas giant flies (Insecta, Diptera, Mydidae), with descriptions of two new species from Central and South America" (PDF). Zootaxa. Auckland, New Zealand: Magnolia Press. 4048 (3): 392 – 411. ISSN 1175-5326. Retrieved 17 March 2017.
- Wilcox, J.; Papavero, N. (1971). "The American genera of Mydidae (Diptera) with the description of three new genera and two new species". Arquivos de Zoologia. São Paulo. 21 (2): 41–119 & 2 Plates. Retrieved 17 March 2017.
- Wilcox, J.; Papavero, N. (1975). "Studies of Mydidae (Diptera) systematics and evolution. III. The genus Messiasia d’Andretta in the Americas (Mydinae)". Arquivos de Zoologia. São Paulo. 26 (1): 1–48. Retrieved 17 March 2017.
- Bequaert, M. 1963. Contribution a la connaissance morphologique et a la classification des Mydaidae (Diptera). Bull. Inst. Roy. Sciences Natur. Belg. 37:1-18.
- Oosterbroek, P. 1998. 41. Mydidae. In P. Oosterbroek: The families of Diptera of the Malay Archipelago. Brill: Boston. p. 95.
- Sack P. (1934) 23. Mydaidae. In: Lindner E. (Ed) Die Fliegen der palaearktischen Region. Band 4(5), 1–29. Schweizerbart‘sche Verlagsbuchhandlung, Stuttgart, 1–29.Keys to Palaearctic species but now needs revision (in German).
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