From Wikipedia, the free encyclopedia
Jump to: navigation, search
Nemastoma bimaculatum from Belgium
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Arachnida
Order: Opiliones
Suborder: Dyspnoi
Superfamily: Nemastomatoidea
Family: Nemastomatidae
Simon, 1872
see text
c. 170 species

The Nemastomatidae are a family of harvestmen with about 170 described species in 16 recent genera. Several fossil species and genera are known.

Unlike some related currently recognized families, the Nemastomatidae are monophyletic.[1]


Members of the Nemastomatidae range in body length from about one to almost six millimeters. Their chelicerae are of normal proportians, but the pedipalps are very elongated and thin in some groups. Leg length is likewise variable.[1]


The subfamily Ortholasmatinae (ten species in two genera) occur on both sides of the Pacific Ocean: in western North America from Mexico to British Columbia and eastern Asia (Japan and northern Thailand. The other subfamily, Nemastomatinae, occur all over Europe up to Iceland and the Caucasus, in the Atlas Mountains of North Africa, from Anatolia to northern Iran, with a few species found outside this region, in Central Asia and the Himalayas. Most species are restricted to very small mountainous southern regions.[1]


The Nemastomatidae are probably a sister group to the genera Dicranolasma (Dicranolasmatidae) and Trogulus (Trogulidae).[1]


The genus name Nemastoma is a combination of Ancient Greek nema "thread" and stoma "mouth", referring to the elongated pedipalps.[1]


Subfamily Nemastomatinae[edit]

  • subgenus Nemastoma
  • Nemastoma bacilliferum bacilliferum Simon, 1879Pyrenees
  • Nemastoma bacilliferum simoni Roewer, 1914
  • Nemastoma bacilliferum simplex Simon, 1913
  • Nemastoma bacilliferum dipentatum Rambla, 1959
  • Nemastoma bidentatum bidentatum Roewer, 1914
  • Nemastoma bidentatum sparsum Gruber & Martens, 1968
  • Nemastoma bidentatum relictum Gruber & Martens, 1968
  • Nemastoma triste triste C. L. Koch, in Herrich-Schaeffer 1835
  • subgenus Dromedostoma
  • Nemastoma acrospinosum acrospinosum Roewer, 1951
  • Nemastoma acrospinosum pretneri Hadzi, 1973
  • subgenus Lugubrostoma
  • Nemastoma lugubre lugubre (Müller, 1776)
  • Nemastoma lugubre bimaculatum (Fabricius, 1775) — Europe
  • Nemastoma lugubre unicolor Roewer, 1914 — southern Europe
  • Paranemastoma quadripunctatum quadripunctatum (Perty, 1833)
  • Paranemastoma quadripunctatum aurosum L. Koch, 1869
  • Paranemastoma quadripunctatum armatum Kulczynski, 1909 — southeastern Europe
  • Paranemastoma quadripunctatum thessalum Simon, 1885 — Greece
  • Paranemastoma quadripunctatum werneri Kulczynski, 1903Asia Minor
  • Paranemastoma quadripunctatum humerale C. L. Koch, 1839 — southeastern Europe
  • Paranemastoma aurigerum aurigerum Roewer, 1951
  • Paranemastoma aurigerum joannae Starega, 1976
  • Carinostoma elegans elegans (Sørensen, 1894)
  • Carinostoma elegans batorligetiense (Szalay, 1951)
  • subgenus Centetostoma
  • subgenus Cretostoma
  • Mitostoma chrysomelas chrysomelas (Hermann, 1804)
  • Mitostoma chrysomelas michielii Hadzi, 1973
  • Mitostoma chrysomelas multidenticulatum Hadzi, 1973
  • Mitostoma chrysomelas alpinum Hadzi, 1931Yugoslavia
  • Mitostoma olgae olgae Silhavý, 1946
  • Mitostoma olgae decorum Silhavý, 1946
  • Mitostoma olgae kratochvili Silhavý, 1946
  • Mitostoma olgae zorae Hadzi, 1973

Subfamily Ortholasmatinae[edit]

incertae sedis[edit]

  • Pyza Starega, 1976


  1. ^ a b c d e Gruber, Jürgen (2007): Nemastomatidae. Simon, 1872. In: Pinto-da-Rocha et al. 2007: 148ff
  2. ^ Giribet & Dunlop 2005


  • Joel Hallan's Biology Catalog: Nemastomatidae
  • Giribet, Gonzalo & Dunlop, Jason A. (2005): First identifiable Mesozoic harvestman (Opiliones: Dyspnoi) from Cretaceous Burmese amber. Proceedings of The Royal Society B 272(1567): 1007-1013. doi:10.1098/rspb.2005.3063 PMID 1599874 - PDF
  • Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds.) (2007): Harvestmen - The Biology of Opiliones. Harvard University Press ISBN 0-674-02343-9