Temporal range: 70 Ma Late Cretaceous - recent
|Nipa palms in Bohol, the Philippines|
Nypa fruticans, commonly known as the nipa palm, is a species of palm native to the coastlines and estuarine habitats of the Indian and Pacific Oceans. It is the only palm considered adapted to the mangrove biome. This species is the only member of the genus Nypa and the subfamily Nypoideae, forming monotypic taxa.
The nipa palm's trunk grows beneath the ground and only the leaves and flower stalk grow upwards above the surface. Thus, it is an unusual palm tree, and the leaves can extend up to 9 m (30 ft) in height. The flowers are a globular inflorescence of female flowers at the tip with catkin-like red or yellow male flowers on the lower branches. The flower produces woody nuts arranged in a globular cluster up to 25 cm (10 in) across on a single stalk. The ripe nuts separate from the ball and are floated away on the tide, occasionally germinating while still water-borne.
Nypa fruticans is also known as attap (Singapore), nipa or sasa (Philippines), buah atap (Indonesia), buah nipah (Malaysia), dừa nước (Vietnam), ging pol (Sri Lanka),jak - จาก (Thailand), gol pata (West Bengal, Bangladesh), and dani (Burma).
- Nypa fruticans Wurmb 
- Nypa fruticans var. neameana F.M.Bailey synonym of Nypa fruticans Wurmb 
Nipa palms grow in soft mud and slow-moving tidal and river waters that bring in nutrients. The palm can be found as far inland as the tide can deposit the floating nuts. It is common on coasts and rivers flowing into the Indian and Pacific Oceans, from Bangladesh to the Pacific Islands. The plant will survive occasional short-term drying of its environment. It is considered native to China (Hainan region), the Ryukyu Islands, Bangladesh, India, Sri Lanka, the Andaman and Nicobar Islands, Cambodia, Thailand, Vietnam, Borneo, Java, Maluku, Malaya, the Philippines, Sulawesi, Sumatra, the Bismarck Archipelago, New Guinea, the Solomon Islands, the Caroline Islands, Queensland, and the Australian Northern Territory. It is reportedly naturalized in Nigeria, the Society Islands of French Polynesia, the Mariana Islands, Panama, and Trinidad.
The long, feathery leaves of the nipa palm are used by local populations as roof material for thatched houses or dwellings. The leaves are also used in many types of basketry and thatching. Large stems are used to train swimmers in Burma as it has buoyancy.
Food and beverages
In the Philippines and Malaysia, the flower cluster (inflorescence) can be tapped before it blooms to yield a sweet, edible sap collected to produce a local alcoholic beverage called tuba, bahal, or tuak. Tuba can be stored in tapayan (balloon vases) for several weeks to make a kind of vinegar known as sukang paombong in the Philippines and cuka nipah in Malaysia. Tuba can also be distilled to make arrack, locally known as lambanog in Filipino and arak in Indonesian.
Young shoots are also edible and the flower petals can be infused to make an aromatic tisane. Attap chee (Chinese: 亞答子; pinyin: yà dá zǐ) (chee meaning "seed" in several Chinese dialects) is a name for the immature fruits—sweet, translucent, gelatinous balls used as a dessert ingredient in Thailand, Malaysia, the Philippines, and Singapore.
The nipa palm produces a very high yield of sugar-rich sap. Fermented into ethanol or butanol, the sap may allow the production of 6,480–15,600 liters (per year)[clarification needed] of fuel per hectare. Sugarcane yields 5,000–8,000 liters per hectare (per year) and an equivalent area planted in corn (maize) would produce just 2000 liters (per year) per hectare, before accounting for the energy costs of the cultivation and alcohol extraction.
While only one species of Nypa now exists, N. fruticans, with a natural distribution extending from Northern Australia, through the Indonesian Archipelago, the Philippine Islands up to China, the genus Nypa once had a nearly global distribution in the Eocene (56-33.4 million years ago).
Fossils of Nypa have also been recovered from throughout the New World, in North and South America, dating from at least the Maastrichtian period of the Cretaceous, through the Eocene making its last appearance in the fossil record of North and South America in the late Eocene.
Assuming the habitat of extinct Nypa is similar to that of the extant species N. fruticans, the presence of Nypa fossils may indicate monsoonal or at least seasonal rainfall regimes, and is likely indicative of tropical climates. The worldwide distribution of Nypa in the Eocene, especially in deposits from polar latitudes, is supporting evidence that the Eocene was a time of global warmth, prior to the formation of modern polar ice-caps at the end of the Eocene.
- The Plant List Nypa fruticans
- John Leslie Dowe. Australian Palms: Biogeography, Ecology and Systematics. p. 83. Retrieved April 20, 2012.
- Flora of China, v 23 p 143, Nypa fruticans
- Wurmb, Friedrich von. 1779. Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen 1: 349, Nypa fruticans
- Kew World Checklist of Selected Plant Families, Nypa fruticans
- "Nypa fruticans". International Union for Conservation of Nature and Natural Resources. Retrieved 2015-10-13.
- "船浦ニッパヤシ群落保護林の保護管理検討委員会報告書" [Funaura nipa palm habitat conservation and management committee report] (PDF). Kyushu Regional Forest Office. p. 6. Retrieved 2015-10-13.
- Gee, Carole T. "The mangrove palm Nypa in the geologic past of the New World." Wetlands Ecology and Management 9.3 (2001): 181-203.
- Singh R. S., 1999, Diversity of Nypa in the Indian subcontinent; Late Cretaceous to Recent. The Palaeobotanist 48(2):147-154.
- Pole, Mike S., and Mike K. Macphail. "Eocene Nypa from Regatta Point, Tasmania." Review of Palaeobotany and Palynology 92.1 (1996): 55-67
- Gee, Carole T. "The mangrove palm Nypa in the geologic past of the New World." Wetlands Ecology and Management 9.3 (2001): 181-203
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