The penis of the Opiliones (harvestmen) is an intromittent organ that is not present in other arachnids. It consists of a long shaft (the truncus) and a terminal capsule called a glans, containing a stylus and ejaculatory duct. It may have from one to three muscles, or none as in the specialized lineage Grassatores, where the penis is operated by haemolymph pressure. In some members of the Cyphophthalmi, the structure has been termed a "spermatopositor".
- Opiliones reproductive anatomy- Dr. Jefferey Schultz, University of Maryland
- Harvestman Spider penis, Thelbunus mirabilus - Australian Museum
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