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Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Infraorder: Araneomorphae
Family: Pacullidae
Simon, 1894
4 genera, 39 species

Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894.[1] It was merged into Tetrablemmidae in 1958,[2][3] then raised back to family status after a large phylogenetic study in 2017.[4]


The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.[4]


Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.[4]

"Lost trachea clade"






Genera and species[edit]

As of April 2019, the World Spider Catalog accepts the following genera and species:[5]


  1. ^ Simon, E. (1894). Histoire naturelle des araignées.
  2. ^ Bonnet, P. (1958). "Bibliographia araneorum". Douladoure, Toulouse. 2 (4): 3297.
  3. ^ Brignoli, P. M. (1973). "Ragni della Melanesia, I. Un nuovo Tetrablemma di Guadalcanal (Isole Salomone) (Araneae Tetrablemmidae)". Memorie della Società Entomologica Italiana, Genova. 72: 86.
  4. ^ a b c Wheeler, Ward C.; et al. (2016), "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling", Cladistics, doi:10.1111/cla.12182
  5. ^ "Family: Pacullidae Simon, 1894". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-04-22.