Papilio memnon, the great Mormon, is a large butterfly native to southern Asia that belongs to the swallowtail family. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.
Its range includes north-eastern India (including Sikkim, Assam and Nagaland), Nepal, Bangladesh, Myanmar, Nicobar Islands, Andaman Islands (stragglers only), western, southern and eastern China (including Hainan), Taiwan southern Japan, Ryukyu Islands, Thailand, Laos, Vietnam, Kampuchea, Malaysia and Indonesia (Sumatra, Mentawai Islands, Nias, Batu, Simeulue, Bangka, Java, Kalimantan and the Lesser Sunda Islands).
Description and polymorphy
The butterfly is large with a 120 to 150 millimetres (4.7 to 5.9 in) span. It has four male and many female forms, the females being highly polymorphic and many of them being mimics of unpalatable butterflies. This species has been studied extensively for understanding the genetic basis for polymorphy and Batesian mimicry. As many as twenty-six female forms are reported.
Typical form agenor
- Male. Tailless, above deep blue to black. It may or may not have red streak on the forewing at the base of the cell.
- Female. Tailless. Upperside forewing ground colour sepia, streaked with greyish white. The basal third part of the cell is red and is touched outwardly with white. Upper hindwing is blue black. It has five to seven yellow or white discal patches.
Female form butlerianus
- Tailless. Resembles the typical male. Both wings are dark sepia. The forewing has a white area on the inner margin. The hindwing is scaled with blue.
Female form alcanor
- Tailed. The sides of the abdomen are yellow.
- Upperside forewing greyish brown with veins and streaks between them black. The cell is red at the base. There is a velvety black patch at the bases of veins 1 and 2 of the upperside forewing.
- Upperside hindwing is black with part of the cell white. There are white streaks around it. The tornus is red with a large black spot. There is a row of red terminal spots between the vein.
Male and female form polymnestoroides
- Male. Upperside hindwing and forewing have short blue discal stripes.
- Female. The upperside forewing is sepia with pale grey streaks amongst the veins. The base of the cell is red. The upperside hindwing is velvety brown with a blue discal area and has black spots, as in the case of the blue Mormon (P. polymnestor), which it mimics.
This species flies up to 2,100 metres (6,900 ft) in the Himalayas, but is most common at low elevations.
This butterfly is found in forest clearings. It is very common and is also seen amongst human habitation. It visits flowers of Poinsettia, Jasminum, Lantana, Canna and Salvia. It usually flies 2 to 4 metres (6 ft 7 in to 13 ft 1 in) above the ground. The butterfly is known to mud-puddle. The males are much more common than females. The female forms butlerianus and alcanor are especially uncommon.
The larva resembles that of the common Mormon (P. polytes), being green with whitish markings. It is heavily parasitised.
- C. A. Clarke, P. M. Sheppard & I. W. B. Thornton (1968). "The genetics of the mimetic butterfly Papilio memnon L." Philosophical Transactions of the Royal Society B. 254 (791): 37–89. Bibcode:1968RSPTB.254...37C. doi:10.1098/rstb.1968.0013. JSTOR 2416804.
- C. A. Clarke & P. M. Sheppard (1971). "Further studies on the genetics of the mimetic butterfly Papilio memnon L." Philosophical Transactions of the Royal Society B. 263 (847): 35–70. Bibcode:1971RSPTB.263...35C. doi:10.1098/rstb.1971.0109. JSTOR 2417186.
- C. A. Clarke & P. M. Sheppard (1973). "The genetics of four new forms of the mimetic butterfly Papilio memnon L.". Proceedings of the Royal Society B: Biological Sciences. 184 (1074): 1–14. Bibcode:1973RSPSB.184....1C. doi:10.1098/rspb.1973.0027. JSTOR 76137. S2CID 86397695.
- Collins, N. Mark; Morris, Michael G. (1985). Threatened Swallowtail Butterflies of the World: The IUCN Red Data Book. Gland & Cambridge: IUCN. ISBN 978-2-88032-603-6 – via Biodiversity Heritage Library.
- Evans, W.H. (1932). The Identification of Indian Butterflies (2nd ed.). Mumbai, India: Bombay Natural History Society.
- Haribal, Meena (1992). The Butterflies of Sikkim Himalaya and Their Natural History. Gangtok, Sikkim, India: Sikkim Nature Conservation Foundation.
- Wynter-Blyth, Mark Alexander (1957). Butterflies of the Indian Region. Bombay, India: Bombay Natural History Society. ISBN 978-8170192329.
- Leech, John Henry (1892–94). Butterflies from China, Japan, and Corea. London: R.H. Porter.
- Media related to Papilio memnon at Wikimedia Commons