European spadefoot toad
|European spadefoot toads|
|Pelobates fuscus fuscus|
|The distribution of extant pelobatids (in black).|
The European spadefoot toads are a family of frogs, the Pelobatidae, with only one extant genus Pelobates, containing four species. They are native to Europe, the Mediterranean, northwestern Africa, and western Asia.
The European spadefoot toads are small- to large-sized frogs, up to 10 cm (3.9 in) in length, which are often inconspicuously coloured. They are predominantly fossorial (burrowing) frogs, which burrow in sandy soils. They have hardened protrusions on their feet to aid in digging, which is the source of the common name. They emerge from the ground during periods of rain and breed in pools, which are usually temporary.
All of the species from this family have free-living, aquatic tadpoles. The eggs are laid in temporary ponds that may quickly evaporate, so the tadpole stage is unusually brief, with rapid development to the adult form in as little as two weeks. To further speed their growth, some of the tadpoles are cannibalistic, eating their brood-mates to increase their supply of protein.
The seven species of American spadefoot toads (genera Scaphiopus and Spea) were previously also included in the family Pelobatidae, but are now generally regarded as the separate family Scaphiopodidae.
- Genus †Elkobatrachus
- †Elkobatrachus brocki
- Genus †Liaobatrachus
- Genus †Eopelobates
- Genus Pelobates
In the Jurassic Morrison Formation, pelobatids are represented by the illium of an unnamed but indeterminate species. This illium is larger than that of Enneabatrachus, a contemporary discoglossid species. A specimen has been recovered from Quarry 9 of Como Bluff in Wyoming. Pelobatids are present in stratigraphic zones 5 and 6 of the formation.
- Foster, J. (2007). "Pelobatidae indet." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. p. 137.
- Zweifel, Richard G. (1998). Cogger, H.G.; Zweifel, R.G. (eds.). Encyclopedia of Reptiles and Amphibians. San Diego: Academic Press. p. 88. ISBN 0-12-178560-2.
- Tron, François (2005). "The Eastern spadefoot Toad (Pelobates syriacus): A new amphibian species for Lebanon" (PDF). Archived from the original (PDF) on 4 August 2014. Cite journal requires
- Henrici, Amy C.; Haynes, Simon R. (2006). "Elkobatrachus brocki, a new Pelobatid (Amphibia: Anura) from the Eocene Elko Formation of Nevada". Annals of Carnegie Museum. 75 (1): 11–35. doi:10.2992/0097-4463(2006)75[11:ebanpa]2.0.co;2.
- Foster, J. (2007). "Appendix." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. pp. 327-329.
- Gissi, Carmela; Diego San Mauro; Graziano Pesole; Rafael Zardoya (February 2006). "Mitochondrial phylogeny of Anura (Amphibia): A case study of congruent phylogenetic reconstruction using amino acid and nucleotide characters" (PDF). Gene. 366 (2): 228–237. doi:10.1016/j.gene.2005.07.034. PMID 16307849. Archived from the original (PDF) on 4 August 2014.
- Roelants, Kim; Franky Bossuyt (February 2005). "Archaeobatrachian paraphyly and pangaean diversification of crown-group frogs" (PDF). Systematic Biology. 54 (1): 111–126. doi:10.1080/10635150590905894. PMID 15805014. Archived from the original (PDF) on 4 August 2014.
- San Mauro, Diego; Miguel Vences; Marina Alcobendas; Rafael Zardoya; Axel Meyer (May 2005). "Initial diversification of living amphibians predated the breakup of Pangaea" (PDF). American Naturalist. 165 (5): 590–599. doi:10.1086/429523. PMID 15795855. Archived from the original (PDF) on 4 August 2014. Retrieved 10 February 2011.
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