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A piebald or pied animal is one that has a spotting pattern of large unpigmented, usually white, areas of hair, feathers, or scales and normally pigmented patches, generally black. The colour of the animal's skin underneath its coat is also pigmented under the dark patches and unpigmented under the white patches. This alternating colour pattern is irregular and asymmetrical. Animals with this pattern may include horses, dogs, birds, cats, pigs, and cattle, as well as snakes such as the ball python. Some animals also exhibit colouration of the irises of the eye that match the surrounding skin (blue eyes for pink skin, brown for dark). The underlying genetic cause is related to a condition known as leucism.
In British English piebald (black and white) and skewbald (white and any colour other than black) are together known as coloured. In North American English, the term for this colouring pattern is pinto, with the specialized term "paint" referring specifically to a breed of horse with American Quarter Horse or Thoroughbred bloodlines in addition to being spotted, whereas pinto refers to a spotted horse of any breed. In American usage, horse enthusiasts usually do not use the term "piebald," but rather describe the colour shade of a pinto literally with terms such as "black and white" for a piebald, "brown and white," or "bay and white," for skewbalds, or color-specific modifiers such as "bay pinto", "sorrel pinto," "buckskin pinto," and such.
Genetically, a piebald horse begins with a black base coat colour, and then the horse also has an allele for one of three basic spotting patterns overlaying the base colour. The most common coloured spotting pattern is called tobiano, and is a dominant gene. Tobiano creates spots that are large and rounded, usually with a somewhat vertical orientation, with white that usually crosses the back of the horse, white on the legs, with the head mostly dark. Three less common spotting genes are the sabino, frame and splash overo genes, which create various patterns that are mostly dark, with jagged spotting, often with a horizontal orientation, white on the head. The frame variant has dark or minimally marked legs. The sabino pattern can be very minimal, usually adding white that runs up the legs onto the belly or flanks, with "lacy" or roaning at the edge of the white, plus white on the head that either extends past the eye, over the chin, or both. The genetics of overo and sabino are not yet fully understood, but they can appear in the offspring of two solid-coloured parents, whereas a tobiano must always have at least one tobiano parent.
The various types of magpie gave their name to pied coloration. The bald eagle derives its name from the word "piebald" in reference to the contrast of its white head and tail with dark body. The term was used in The Conservationist by Nadine Gordimer when referring to kingfishers.
Many other animal species may also be "pied" or piebald including, but not limited to, squirrels and birds. Snakes, especially ball pythons and corn snakes, may also exhibit seemingly varying patches of completely pigmentless scales along with patches of pigmented scales. Some domesticated foxes born from the Russian Institute of Cytology and Genetics also carry this coloring. Recently, a piebald blood python was discovered in Sumatra.
Bicolor cats carry the piebald gene. The same pattern that applies to cats also applies to dogs when the white spotting gene involved is indeed piebald and not another white-causing gene found in dogs. The piebald gene is also found in domestic guinea pigs, goldfish, rabbits, hamsters, goats, fancy rats, cows, and ferrets.
- Heterochromia iridum
- Harper, Douglas. Online Etymology Dictionary.
- Skeat, Walter W. (1882). The Concise Dictionary of English Etymology. Hertfordshire: Clarendon Press. p. 442.
- "Introduction to Coat Color Genetics" from Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web Site accessed January 12, 2008
|Wikisource has the text of the 1911 Encyclopædia Britannica article Pie.|
- "Piebald Moose". Ontario Out of Doors . 17 March 2012. Retrieved 15 December 2014. (extremely rare wild piebald moose photographed in northwest Alberta, Canada)