Protungulatum donnae

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Temporal range: Late Cretaceous - Paleocene, Maastrichtian–Danian

Possible Campanian record

Protungulatum donnae.JPG
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Genus: Protungulatum
Sloan and Van Valen, 1965
Species: P. donnae

Protungulatum donnae is the type species for the Protungulatum genus, an extinct early form of eutherian mammal. Though it is by no means the earliest mammal in the fossil record, a 2013 study considers P. donnae to be the oldest undisputed placental mammal fossil,[1][2] though more recent examinations conclude that it was a more basal eutherian and that no placental predates the Paleocene.[3]


  1. ^ O'Leary, Maureen A.; Bloch, Jonathan I.; Flynn, John J.; Gaudin, Timothy J.; Giallombardo, Andres; Giannini, Norberto P.; Goldberg, Suzann L.; Kraatz, Brian P.; Luo, Zhe-Xi; Meng, Jin; Novacek, Michael J.; Perini, Fernando A.; Randall, Zachary S.; Rougier, Guillermo; Sargis, Eric J.; Silcox, Mary T.; Simmons, Nancy b.; Spaulding, Micelle; Velazco, Paul M.; Weksler, Marcelo; Wible, John r.; Cirranello, Andrea L. (8 February 2013). "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667. Bibcode:2013Sci...339..662O. doi:10.1126/science.1229237. PMID 23393258. Retrieved 9 February 2013. 
  2. ^ Wilford, John Noble (7 February 2013). "Rat-Size Ancestor Said to Link Man and Beast". New York Times. Retrieved 9 February 2013. 
  3. ^ Resolving the relationships of Paleocene placental mammals