von Meyer, 1847
Psephophorus is an extinct genus of sea turtle that lived from the Oligocene to the Pliocene. Its remains have been found in Europe, Africa, North America, and New Zealand. It was first named by Hermann von Meyer in 1847, and contains seven species, P. polygonus, P. calvertensis, P. eocaenus, P. oregonesis, P. californiensis, P. rupeliensis, P. scaldii, and a species discovered in 1995, P. terrypratchetti.
Discovery and identification
Von Meyer originally named Psephophorus in 1846. At first he was unable to identify the creature beyond its dermal plates, but when he later received a drawing he was able to describe the specimen, which was then in Pressburg, as a fragment of a carapace, which contained seventy bones.
In 1879, H. G. Seeley was asked to study the Psephophorus specimen by Franz Ritter von Hauer, the Director of the Austro-Hungarian Imperial and Royal Geological Survey. Up until then, the specimen's identity had been undeterminable, with even Seeley describing it at first to seem like "the dermal covering of an Edentate closely allied to the Armadilloes." Seeley examined some bone fragments and concluded the specimen was that of a reptilian creature, furthermore a chelonid. It also proved to be more closely related to Sphargis than any other type in the Chelonian order.
Relation to modern Leatherback sea turtles
For a long time, modern Leatherback sea turtles (Dermochelys) were believed to be descended directly from Psephophorus, specifically the species P. polygonus. However, a 1996 analysis by Wood et al. proved that most of the taxa in the two genera were not connected, meaning Psephophorus could not be a direct ancestor of the modern leatherbacks. The platelets on Psephophorus are quite similar to those on Dermochelys, despite differences in outer morphology and size. The platelet comprises an external compact layer and an internal zone of cancellous bone.
- Psephophorus polygonus is the type species, and was discovered by von Meyer in 1846. Fossils of the species have shown bony, carapacial ridges, adjacent, small and polygonal bony ossicles which measure about 22 × 25 mm. When the ossicles extend further along the longitudinal axis, their size increases up to an average of 33 × 41 mm.
- Psephophorus calvertensis was first named by Palmer in 1909. It was so named for being found in the Calvert Formation. P. calvertensis is a rarely found specimen.
- Psephophorus eocaenus was first named by Andrews in 1901.
- Psephophorus californiensis was first named by Gilmore in 1937.
- In 1990s, Richard Köhler found a specimen of the genus Psephophorus in New Zealand, which proved to be different from the species which had been found in Europe, Egypt, and the United States. It was then declared a new species altogether, and Köhler named the species Psephophorus terrypratchetti after the author Terry Pratchett, who wrote a series of fantasy books set on a world carried on the back of a giant turtle.[unreliable source?]
- Kohler, R. (September 1995). "Köhler - A new species of Psephophorus". Journal of the Royal Society of New Zealand. 25 (3): 371–384. doi:10.1080/03014223.1995.9517495.
- "The Oregon Fossil Guy". www.mailtribune.com. 2008-04-27. Retrieved 2008-06-28.
- "Sharktooth Hill Fauna, circa 2003". Shark Tooth Hill.com (optionally viewed as a Microsoft Word document). 2003. Archived from the original on 2012-12-09. Retrieved 2008-06-28.
- Proceedings; page 8. By the Zoological Society of London; published 1891. Retrieved on June 28th, 2008.
- "New Zealand species of Psephophorus". Everything2.com (partially derived from an interview with Terry Pratchett on The Discworld Companion). 2001-11-01. Retrieved 2008-06-27.
- "New Remains of Psephophorus polygonus (Chelonii: Dermochelyidae) from the Miocene of Southern Italy" (PDF). digilander.libero.it. 2004. Retrieved 2008-06-27.
- A Manual of Palaeontology; page 1091. By Henry Alleyne Nicholson and Richard Lydekker; published 1889, Blackwood. Retrieved on June 28th, 2008.
- Seeley, H. G. (1880). "Note on Psephophorus polygonus, v. Meyer, a new Type of Chelonian Reptile allied to the Leathery Turtle". Quarterly Journal of the Geological Society. 36 (406–413): 406–413. doi:10.1144/GSL.JGS.1880.036.01-04.32.
- "Turtles of the World". nlbif.eti.uva.nl. Archived from the original on 2011-06-08. Retrieved 2008-06-27.
- "Fossil sea turtles (Chelonii, Dermochelyidae and Cheloniidae) from the Miocene of Pietra Leccese (late Burdigalian-early Messinian), Southern Italy" (PDF). 2007. Archived from the original (PDF) on 2009-04-19. Retrieved 2008-06-27.
- "Bone histological results of Testudinata" (PDF). hss.ulb.uni-bonn.de. Retrieved 2008-06-27.[dead link]
- "Psephophorus calvertensis at the Paleobiology Database". paleodb.org. Retrieved 2008-06-27.
- Weems, Robert E. (1974). "Middle Miocene sea turtles (Syllomus, Procolpochelys, Psephophorus) from the Calvert Formation". Journal of Paleontology. 48 (2): 279–303.
- "Psephophorus eocaenus at the Paleobiology Database". paleodb.org. Retrieved 2008-06-27.
- Curiosities of Biological Nomenclature
- Callaway, Jack M.; Nicholls, Elizabeth L. Ancient Marine Reptiles.
- Nielsen, Eigil (1959). "Eocene Turtles from Denmark" (PDF). Bulletin of the Geological Society of Denmark. 14 (2): 96–114.
- Wieland, G. R. (1900). "Some Observations on Certain Well-Marked Stages in the Evolution of the Testudinate Humerus". American Journal of Science. Series 4. 9 (54): 413–424 [p. 421]. doi:10.2475/ajs.s4-9.54.413.
- Hay, O. P. (1898). "On Protostega, the Systematic Position of Dermochelys, and the Morphogeny of the Chelonian Carapace and Plastron". The American Naturalist. 32 (384): 929–948. doi:10.1086/277069.