Pterodactyloidea

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Pterodactyloids
Temporal range:
Late Jurassic - Late Cretaceous, 162.7–66 Ma
Pterodactylus antiquus - IMG 0681.jpg
Cast of a Pterodactylus antiquus specimen, Carnegie Museum of Natural History
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Pterosauria
Clade: Caelidracones
Suborder: Pterodactyloidea
Plieninger, 1901
Subgroups
Synonyms

Dracochira Haeckel, 1895

Pterodactyloidea (derived from the Greek words πτερόν (pterón, for usual ptéryx) "wing", and δάκτυλος (dáctylos) "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoidea by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth. Many species had well developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids (specifically the family Azhdarchidae) were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles.

"Pterodactyl" is also a common term for pterodactyloid pterosaurs, though it can also be used to refer to Pterodactylus specifically or (incorrectly) to pterosaurs in general. Well-known examples of pterodactyloids include Pterodactylus, Dsungaripterus, Pteranodon, and Quetzalcoatlus.

In 2014, fossils from the Shishugou Formation of China were classified as the most basal pterodactyloid yet found, Kryptodrakon. It is about 5 million years older than the oldest previously known confirmed specimens.[1] Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,[2] though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur.[1]

Classification[edit]

Pterodactyloidea is traditionally considered to be the group of short-tailed pterosaurs with long wrists (metacarpus), compared with the relatively long tails and short wrist bones of basal pterosaurs ("rhamphorhynchoids"). In 2004, Kevin Padian formally defined Pterodactyloidea as an apomorphy-based clade containing those species possessing a metacarpal at least 80% of the length of the humerus, homologous with that of Pterodactylus.

A subgroup of pterodactyloids, called the Lophocratia, was named by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of Pterodaustro guinazui and Quetzalcoatlus northropi, and all its descendants.[3] This group was named for the presence of a head crest in most known species, though this feature has since been found in more primitive pterosaurs and was probably an ancestral feature for all pterodactyloids.[4]

There are competing theories of pterodactyloid phylogeny. Below is a cladogram showing the results of a phylogenetic analysis presented by Andres, Clark & Xu, 2014. This study found the two traditional groupings of ctenochasmatoids and kin as an early branching group, with all other pterodactyloids grouped into the Eupterodactyloidea.[1]

 Caelidracones 

AnurognathidaeAnurognathusDB white background.jpg


 Pterodactyloidea 

Kryptodrakon


Lophocratia
Archaeopterodactyloidea 

GermanodactylidaeGerm rhamph DB.jpg


 Euctenochasmatia 

PterodactylusPterodactylus holotype fly mmartyniuk white background.png




Ardeadactylus


 Ctenochasmatoidea 

Gallodactylidae



Ctenochasmatidae






Eupterodactyloidea 

Haopterus


 Ornithocheiroidea 
 Pteranodontia 

NyctosauridaeNyctosaurus DB white background.jpg


 Pteranodontoidea 

PteranodonPteranodon longiceps mmartyniuk wiki.png




Hongshanopterus




Lonchodectes




Istiodactylidae




Boreopteridae




Cearadactylus




Brasileodactylus




Ludodactylus




AnhangueridaeLiaoningopterusDB flipped.jpg



 OrnithocheiridaeOrnithocheirus BW flipped.jpg












 Azhdarchoidea 


Bennettazhia




TapejaridaeBakonydraco1DB.jpg




 Neoazhdarchia 


Chaoyangopteridae



AzhdarchidaeQuetzalcoatlus07.jpg





ThalassodrominaeTupux longDB2.jpg



Dsungaripteridae










Some studies based on a different type of analysis have found that this basic division into primitive (archaeopterodactyloid) and advanced (eupterodactyloid) species may not be correct. In 2014, Steven Vidovic and David Martill concluded that several pterosaurs traditionally thought of as archaeopterodactyloids, specifically Gladocephaloideus, Cycnorhamphus, Aurorazhdarcho, Aerodactylus, and Ardeadactylus, may actually have been more closely related to ornthocheiroids and azhdarchoids. They found that Pterodactylus itself was actually more primitive than both this new group, the Aurorazhdarchidae, and the classic archaeopterodactyloid ctenochasmatids. The results of their analysis are shown below.[5]

Pterodactyloidea

Pterodactylus antiquusPterodactylus holotype fly mmartyniuk white background.png




Pterodactylus kochiPterodactylus holotype fly mmartyniuk white background.png


Lophocratia

Ctenochasmatidae





Germanodactylus rhamphastinusGerm rhamph DB.jpg


Aurorazhdarchidae


Aerodactylus scolopacicepsAerodactylus MCZ 1505.png



Gladocephaloideus jingangshanensis





Cycnorhamphus suevicus




Aurorazhdarcho micronyx



Ardeadactylus longicollum






Eupterodactyloidea
Pteranodontia

Istiodactylidae




Ornithocheirus mesembrinus


Pteranodontoidea


Anhanguera piscator



Ludodactylus sibbicki





Nyctosaurus gracilisNyctosaurus DB white background.jpg



Pteranodon longicepsPteranodon longiceps mmartyniuk wiki.png






Dsungaripteroidea

Germanodactylus cristatus


Tapejaroidea

Dsungaripterus weii


Neoazhdarchia

TupuxuaraTupux longDB2.jpg


Azhdarchoidea

AzhdarchidaeQuetzalcoatlus07.jpg



TapejaridaeBakonydraco1DB.jpg











References[edit]

  1. ^ a b c Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. doi:10.1016/j.cub.2014.03.030. 
  2. ^ Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." Geological Magazine, (advance online publication) doi:10.1017/S0016756811001154
  3. ^ Unwin, D. M., (2003). "On the phylogeny and evolutionary history of pterosaurs." Pp. 139-190. in Buffetaut, E. & Mazin, J.-M., (eds.) (2003). Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217, London, 1-347.
  4. ^ Witton, Mark (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. ISBN 978-0691150611. 
  5. ^ Vidovic, S. U.; Martill, D. M. (2014). "Pterodactylus scolopaciceps Meyer, 1860 (Pterosauria, Pterodactyloidea) from the Upper Jurassic of Bavaria, Germany: The Problem of Cryptic Pterosaur Taxa in Early Ontogeny". PLoS ONE. 9 (10): e110646. doi:10.1371/journal.pone.0110646.