|Copris lunaris (Coprini) ♀|
About one dozen; see text
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.
Scarabaeinae tribes, with some selected genera and species, are:
- Gymnopleurini (sometimes included in Scarabaeini)
- Phanaeini MacLeay, 1819
- Sisyphini (sometimes included in Scarabaeini)
- Unknown tribe / Incertae sedis
Coprini, Oniticellini, Onitini, Onthophagini and Phanaeini are sometimes separated as subfamily Coprinae.
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