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Temporal range: Late Triassic, Carnian
Scleromochlus BW.jpg
Life restoration of Scleromochlus taylori
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Ornithodira
Clade: Pterosauromorpha
Family: Scleromochlidae
Huene, 1914
Genus: Scleromochlus
Woodward, 1907
Species: S. taylori
Binomial name
Scleromochlus taylori
Woodward, 1907

Scleromochlus (Greek for "hard fulcrum") is an extinct genus of small avemetatarsalians from the Late Triassic period.


Scleromochlus taylori was about 181 mm (about 7.1 inches) long, with long hind legs; it may have been capable of four-legged and two-legged locomotion. Most recent studies about its gait suggest that it engaged in kangaroo- or springhare-like plantigrade hopping;[1][2][3] if Scleromochlus is indeed related to pterosaurs, this may offer insight as to how the latter evolved, since early pterosaurs also show adaptations for saltatorial locomotion.[4]


Its fossils have been found in the Carnian Lossiemouth Sandstone of Scotland. The holotype is BMNH R3556, a partial skeleton preserved as an impression in sandstone; part of the skull and tail are missing.[5]

Scleromochlus is a monotypic genus (single species), including the type species S. taylori.


Skeletal restoration

A lightly built cursorial animal, its phylogenetic position has been debated; as different analyses have found it to be either the basal-most ornithodiran, the sister-taxon to Pterosauria, or a basal member of Avemetatarsalia that lies outside of Ornithodira. In the phylogenetic analyses conducted by Nesbitt et al. (2017) Scleromochlus was recovered either as a basal member of Dinosauromorpha or as a non-aphanosaurian, non-pterosaur early avemetatarsalian. However, the authors stressed that scoring Scleromochlus was challenging given the small size and poor preservation of the fossils, and stated that it could not be scored for many of the important characters that optimize near the base of Avemetatarsalia.


  1. ^ Sereno et Arcucci 1993; 1994
  2. ^ Benton 1999
  3. ^ Witton, Mark P. (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. ISBN 0691150613. 
  4. ^ Witton, Mark P. (2015). "Were early pterosaurs inept terrestrial locomotors?". PeerJ. 3: e1018. PMC 4476129Freely accessible. PMID 26157605. doi:10.7717/peerj.1018. 
  5. ^ Basic information from Fossilsmith