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Scutalus mariopenai 5.png
A live individual of Scutalus mariopenai
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
(unranked): clade Heterobranchia

clade Euthyneura
clade Panpulmonata
clade Eupulmonata
clade Stylommatophora
informal group Sigmurethra

Superfamily: Orthalicoidea
Family: Bulimulidae
Subfamily: Peltellinae[1]
Genus: Scutalus
Albers, 1850[2]

Scutalus is a genus of air-breathing land snails, a terrestrial pulmonate gastropod mollusks in the family Bulimulidae.[1]

Previously this genus was placed within the subfamily Bulimulinae.[3]

Shells of Scutalus sp. on a Moche pot, 200 AD. Larco Museum Collection, Lima, Peru.


Species in the genus Scutalus include:

subgenus Scutalus Albers, 1850

subgenus Vermiculatus Breure, 1978[5]

subgenus ?


  1. ^ a b c Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. doi:10.1127/arch.moll/1869-0963/141/001-020.
  2. ^ Albers J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers: 1-262. Berlin. page 160.
  3. ^ Breure A. S. H. (1979). "Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca)". Zoologische Verhandelingen Leiden 168: 1-215, figs 1-182, pls. 1-3, tables 1-5. PDF.
  4. ^ a b c d e f Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM.
  5. ^ Breure A. S. H. (1978). "Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda)". Zoologische Verhandelingen Leiden 164: 1-255. page 166.
  6. ^ [1]
  7. ^ [2]

External links[edit]

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