|First appearance||Fantastic Four #2 (January 1962)|
Amazing Adult Fantasy #7 (December 1961) (by retcon)
|Created by||Stan Lee (writer)|
Jack Kirby (artist)
|Place of origin||Skrullos; later moved capital to Tarnax IV (destroyed)|
|Pantheon||Kly'bn, Sl'gur't and Zorr'Kiri|
|Notable members||See Known Skrulls|
|Inherent abilities||Advanced shapeshifting|
The Skrulls are a fictional race of extraterrestrial shapeshifters appearing in American comic books published by Marvel Comics. The Skrulls first appeared in Fantastic Four #2 and were created by Stan Lee and Jack Kirby. They originated from the planet Skrullos and their empire is located in the Andromeda Galaxy.
This section needs expansion. You can help by adding to it. (January 2019)
The Skrulls first appeared in Fantastic Four #2 and were created by Stan Lee and Jack Kirby. The Skrulls next appear in Fantastic Four #18. The four who impersonated the members of the Fantastic Four next appear in The Avengers #92. The concept of the Skrulls would be explored and used throughout multiple storylines, spin-off comics, and derivative projects.
Fictional species biography
Millions of years ago, in the main Marvel continuity, the Celestials perform genetic experiments on the reptilian ancestors of the Skrulls, resulting in three branches of Skrulls: The Prime, the Deviants and the Eternals. Eventually the three branches battle with one another, and the Deviant branch – due to possessing the innate ability to shapeshift – were triumphant, and wiped out all members of the other two races until only two were left: the Skrull Eternal, Kly'bn, and Prime Skrull of the original non-Deviant Skrull race. Kly'bn implored them to spare his life, as killing him would kill part of their heritage. The Deviants' leader, Sl'gur't, eventually fell in love with Kly'bn, with the two eventually becoming the gods of the Skrull pantheon. Taking the name Prime Skrull, the last original Skrull escaped to Earth in the 20th century and later became a member of the Underground Legion. From that point, the Skrulls began to expand their territory. The Deviant branch later split into two more groups, the modern Skrulls and an anomaly called the Dire Wraiths.
The Skrulls originate from the planet of Skrullos, and were originally a mercantile civilization, primarily interested in free trade and willing to share their technology with all races they deemed worthy. When they encountered a new race, they simply transformed themselves to resemble that race. The Skrull empire that resulted from these contacts was based on free trade and mutual cooperation.
Eventually, the Skrulls developed long-distance space travel; a great tour of the universe was undertaken, led by Emperor Dorrek. Finally, the Skrull delegation reached the planet Hala, home to the then-barbaric Kree and the peaceful Cotati, and held a contest to determine which of the races would represent Hala to the Skrull Empire. Seventeen members of each race were taken to different uninhabited planetoids where they were left with sufficient supplies for one year. At the end of that period, whichever group had done the most with themselves would be adjudged the most worthy. The Kree were taken to Earth's Moon where they built a great city while the Cotati were taken to another barren world in a different solar system and used their abilities to create a beautiful park. Realizing that the Cotati were going to win, the enraged Kree killed all the Cotati. When the Kree revealed that they had solved the question of who would represent Hala to the Skrulls by destroying their opposition, the Skrull delegates were appalled and vowed that Hala would forever be banned from their circle of favored worlds—so the Kree massacred them as well, took over the Skrull landing spaceship by force, and developed their own technology from it.
Because of the immense distances involved, decades passed before the Skrulls learned of the Kree's activities. By this time it was too late. The Kree were now advanced and audacious enough to attack the Skrulls in their home galaxy. During the millennia that followed, the Kree aggression forced the Skrulls to become a militaristic civilization, and the Skrulls eventually developed the vicious streak needed to conduct intergalactic war. Their entire culture was remade in the warrior image. The Kree–Skrull War continued for thousands of years with only brief interruptions. The Skrull Throneworld is later moved from Skrullos to the more central Tarnax IV, with their empire encompassing over fifty worlds. The Skrulls also created the first Cosmic Cube, which later became the Shaper of Worlds; the Shaper related that, upon attaining self-awareness, he lashed out blindly, destroying two thirds of the Skrull empire and casting the civilization back into barbarism (suggesting this occurred before the war with the Kree broke out).
The Kree Accuser Ronan had wrested control of the Kree Empire from the Supreme Intelligence and attacked the Skrulls, reigniting hostilities. The Avengers become involved once the Super Skrull kidnaps Captain Marvel, and in turn they battle a Kree Sentry robot, three of the original Skrulls that had fought the Fantastic Four, S.H.I.E.L.D. agents, rogue Inhumans and legions of Skrulls. The Supreme Intelligence ends the immediate conflict by revealing to Avengers' ally Rick Jones that he has vast mental potential, which is then used to freeze all combatants in place. The Supreme Intelligence announces that, while the Kree and Skrulls have reached genetic dead ends, the human race displays incredible untapped potential. It is revealed years later that the conflict produced a Kree-Skrull hybrid, Hulkling, who is the child of one-time lovers Captain Marvel and the Skrull Princess Anelle.
Empire changing events
The Skrulls became aware of Earth when they discovered a space warp linking Earth and Throneworld. Later they discovered the Earth possesses an equidistant link to Hala. They see Earth as a world to conquer but they are content to hold off a full invasion. Because of their caution, the Skrulls have sent only a handful of scoutwarriors to infiltrate Earth, but fail to take into account the presence of the modern superhero.
In 1958, Skrulls attempted to sabotage Earth's space program. They battled 3-D Man, and set the Cold Warrior against 3-D Man. Later a small scouting party used their shape shifting powers to impersonate the Fantastic Four, committing crimes so the country would turn against the Fantastic Four, the only ones with the power to stop them. But the Fantastic Four were able to locate the Skrulls and tricked their leaders into believing the Earth was full of threats. The fleet left and Mr. Fantastic made the Skrulls that were left behind shapeshift into cows and he hypnotized them to remember nothing about their true heritage.
In retaliation the Skrull Emperor Dorrek VII dispatches Kl'rt, a Skrull known as the Super-Skrull, to Earth to defeat the Fantastic Four. Kl'rt possesses the powers of the entire Fantastic Four (in addition to shapeshifting and hypnotism), and he holds the team at bay until Mister Fantastic discovers the source of his power and uses a miniature device to jam the ray, before the Human Torch imprisons him inside a crater. The Super-Skrull posed as the jailed Dr. Franklin Storm, after transporting him to the Skrull throneworld, and battled the Fantastic Four as the Invincible Man, another Skrull, ultimately caused the death of Dr. Storm by attaching a bomb to his chest before he was transported back, although Storm turned to the floor, saving the Fantastic Four. The Fantastic Four later travel to Tarnax IV and find the Skrull responsible for the murder of Susan and Johnny Storm's father (which turned out to be Warlord Morrat). Skrulls were involved in other events such as sending the Super-Skrull to battle Captain Mar-Vell, and abducting the Thing as a contestant in the Skrull Games.
Later the cosmic being the Watcher adjudicates a duel between champions from both races: Colonel Bel-Dann of the Kree and Warlord Raksor of the Skrulls. After one year of conflict the duel remains deadlocked and is ultimately abandoned with no resolution. Years later during a conflict with Xandar that included the Fantastic Four and the Champions of Xandar, Emperor Dorrek VII's wife, Empress R'klll, stages a coup and kills him to become ruling Empress.
Empress R'klll's reign is short-lived, however, as the greatest blow to the Skrulls arrives in the form of the World Devourer Galactus. After his Herald Nova annihilates the Skrull fleet, Galactus devours Tarnax IV, killing billions of Skrulls, including R'klll and the Princess Anelle. With the destruction of the central government, the Skrull Empire has collapsed into hundreds of bickering factions. Their galaxy is filled with Governors who have declared themselves to be Emperor of the Skrulls and soon a civil war began. The Shi'ar maintain a heavy presence in the former empire as well, constantly dealing with packets of resistance.
Second Kree-Skrull War
Later Reed Richards discovered that yet another Governor-turned-Emperor, Zabyk, had created a genetics bomb, which, when it exploded made all the Skrulls lose their shape-shifting abilities. Whatever form they were in they were stuck in at the time of the explosion.
In desperation, since without their shapeshifting powers they were vulnerable, they managed to kidnap the herald of Galactus, Nova, in an attempt to get Galactus to destroy Kree worlds for them. Their plot failed when the Silver Surfer helped Galactus rescue Nova in exchange for his freedom from Earth.
When the Celestials made an appearance, this scared the Skrulls into starting a second war with the Kree to show they were not powerless. The attack was first made by Kylor, one of five governors claiming to be Emperor. He had a spy in the Kree Empire, Nenora, who gave him the early edge. They tried to hide their secret weakness at all costs from the Kree.
Kylor was eventually betrayed by Nenora, as was the entire Skrull race. She assumed power of the empire and wished to keep it for herself. It was Empress S'byll who stepped up to control the Skrulls and finish the war. She was able to resurrect the Super Skrull and he returned to find the empire in shambles. They tricked the Silver Surfer into attacking a Badoon fleet, which he was not aware were allied with the Kree. Since the Kree waged war on the Surfer, he allied himself with the Skrulls to save Zenn-La from destruction.
To restore their powers, the Super Skrulls DNA needed to be transferred to S'byll, because only a female could spread the ability to the other Skrulls. The Surfer helped them to power a machine that would return the Empress' shape shifting abilities. The device was successful. With this power she was able to restore every Skrull she touched and became the true Empress of the Skrulls.
She and Super Skrull waged war with the Kree with the help of the Silver Surfer. She replaced Super Skrull with Captain Reptyl and he helped to change her into an emotionless reptile. But Reptyl was betrayed by one of his own and the Skrull fleet was utterly destroyed by the Kree. S'byll and Silver Surfer went alone to attack the Kree homeworld of Hala.
Gaining the knowledge that Nenora was actually a Skrull gave them the advantage. S'byll gave her shape-shifting abilities back to her, crippling her and showing the Kree her true self. A peace treaty was created.
The Skrulls eventually broke that peace treaty as they kept their military drive and prefer to fight rather than negotiate. Among the other races contained within the Skrull Empire are the Druff, Guna, Kallusina, Morani, Pheragot, Queega, Tekton, and Yirbek races. Other races remain independent, whether through treaties or open resistance. Such races include the Clegrimites, Gegku, Krylorians, Wilameanis, and Xandarians. The Skrull-Xandarian war with the Xandarians recently ended with the destruction of Xandar by the supervillainess Nebula.
Skrulls are meanwhile involved in numerous other activities. They were revealed to have conquered a medieval planet and abandoned it after its inhabitants perished. Skrull Prime Ten battled the Fantastic Four and Captain Mar-Vell. The Skrulls sent their own representative to witness the fate of the Phoenix, at her trial by the Shi'ar. There is a lone appearance of Queen Akilll of the Skrull. A lone Skrull poisoned Vera Gantor to force the Avengers to seek the Resurrection Stone on his behalf. The Skrulls sent their General Zedrao to give the dying Captain Mar-Vell the Royal Skrull Medal of Honor. The Skrulls had captured a Kree spy who had infiltrated into the Skrulls by using a symbiote.
A group of genetically-bred Skrulls called Warskrulls began infiltrating the Shi'ar Empire. They used a Nexus Amplifier to assume total physical discorporation of their targets, adopting their super powers along with their physical appearance. They captured and replaced the Starjammers with members of their race as well as key components in government. The Warskrulls, acting under the direction of the paramount among them, posing as a Shi'ar called the Chamberlain, began committing murders and even genocide upon various Shi'ar affiliated races, such as the P!ndyr, in the name of the Shi'ar Empire. One Warskrull replaced the Imperial Consort, Professor X, called himself Warlord, and took mental control over Lilandra, the Majestrix of the Shi'ar. They were eventually defeated by the combined forces of Deathbird and the X-Men and later the Shi'ar vowed to weed out any other Warskrulls.
Some time later a Skrull called Paibok mounted a campaign to invade Earth, using another Skrull called Lyja as an infiltrator. The plan involved secretly replacing the real Alicia Masters, The Thing's blind lover. The plan fell through when she instead seduced Human Torch and the two eventually married one another. The long running facade eventually came to light and the group discovered that the real, human Alicia was actually being held captive by the Skrull nation and Lyja defected, having fallen in love with the Human Torch. The Fantastic Four rescued her, but Lyja seemingly died in the process as she sacrificed herself to save Johnny when Paibok attacked.
Paibok later revealed to Devos the Devastator that she was still alive. Paibok, with the assistance of Devos, awoke Lyja from her comatose state and gave her the power to fly and fire energy blasts from her hands (she called herself "Lyja the Lazerfist"). These powers were later revealed to come from a special device implanted in her. Joining them in search of vengeance, the three arrived on Earth, and battled the Torch at Empire State University. The Torch panicked, used his nova flame and destroyed the campus. Alongside Devos and Paibok, Lyja witnessed a battle between the Fantastic Four and an alternate Fantastic Four. However, Lyja still bore feelings for the Torch and once again betrayed her cohorts to side with the Fantastic Four. This ended with the Fantastic Four destroying the "Skrull War World", the Skrulls' largest attack ship.
A group of Skrulls traveled to Earth where they began plotting to invade the planet once again. They were however at some point approached by Apocalypse who wanted to lure together twelve powerful mutants that he needs to ascend to godhood. At the same time, the body of the Living Monolith, who had been hurled out into the far reaches of space by Thor, is found and brought back to Earth. On Earth, a mutant Skrull named Fiz snuck into Xavier's mansion to tell them urgent news of trouble: the Skrulls have allied themselves with someone powerful and the only thing he knows is that there are twelve mutants that can't be harmed and that something terrible is going to happen. Fiz also reveals the existence of a subtype of Skrulls that possess innate powers like Earth's mutants. Deemed outcasts by the other Skrulls, they were hunted by the Empire.
After the eventual defeat of Apocalypse, Professor X went into space to train them, giving them the name Cadre K. Cadre K and Xavier returned to Earth during the Ruul's attempt to overtake the planet by turning it into an intergalactic prison. A wounded Z'Cann attempted to pass on vital information to Rogue. Their physical contact affected Rogue's powers for some time. The information was received and the X-Men were reunited with Bishop.
The Skrull Empire is the first of the major interstellar empires to be invaded by the forces of Annihilus. The Annihilation Wave's superweapon, the Harvester of Sorrow, physically destroys dozens of Skrull planets. In the aftermath, the Skrulls unsuccessfully attempt to convince Hulkling to become their new Emperor.
The Rise and Fall of the Shi'ar Empire
A group of Warskrulls attacked a remote Shi'ar space station and took it over. Disguised as Shi'ar, they were taken aboard the space station willingly and slaughtered all those aboard except a few to keep working the ship. The group, however was eventually defeated by a group of X-Men (Darwin, Havok, Marvel Girl, Nightcrawler, Professor X, Warpath). These Warskrulls are however revealed to be cyborgs.
In the "Secret Invasion" series by Brian Michael Bendis and Leinil Yu, a flailing Skrull empire, under the leadership of newcomer Queen Veranke, made an all-out effort to infiltrate and conquer Earth. It was revealed that the Skrulls had infiltrated Earth for quite some time by capturing and replacing many of Earth's heroes. They followed this infiltration with their main assault group. This wave of Skrulls formed a new version of the Super-Skrull, with each possessing the abilities of several different heroes. Their invasion ultimately failed, costing the life of Veranke, and reducing their numbers even further. Even their gods were slain, and the Japanese God of Evil Amatsu-Mikaboshi took over their Pantheon, leading to the events of Chaos War.
Reestablishing the Empire and Infinity
After the events of Secret Invasion, the Skrulls, now in smaller numbers, became very segmented with various factions rising up around warlords that began plotting to rise back to power by orchestrating events for their benefits.
During the "Infinity" storyline, several planets that the Skrulls settled on began being targeted by the Builders and while the Silver Surfer tried to save most of them, the Skrulls' population was once more decimated. Kl'rt later represented the Skrull Empire as a member of the Galactic Council. After the Avengers' victory over the Builders and Thanos, Kl'rt becomes the new Emperor of the Skrulls where they settle on Tarnax II.
Meanwhile, Cadre K had settled on a ranch in South Dakota, while another group of Skrulls remain undercover on Earth as the "Warner" family, who are gathering data for the next Skrull invasion. The Warner family participate in day-to-day life, but end up stuck on Earth completely cut off from the empire.
During the search of the Infinity Stones in the "Infinity Countdown" storyline, Razor reveals the true origin of the Tree of Shadows, according to Razor the Tree of Shadows was created eons ago by the Elders of the Universe known as The Gardener who tried to bring life to the Null Space and planted it in the Darkforce Dimension. Eventually Null Space and the Tree of Shadows were discovered by a more pre-historic version of the Shi'ar and the Skrulls, a find that erupted in a war between the two alien races. The war ranged until one member of the Shi'ar was able to grasp one of the seeds of the Tree. His physiology, fear and belief's shaped the birth of the most vicious killing machine the universe had ever seen, the first Raptor and while the Skrulls, being shape-shifters, were still able to hold out against the Shi'ar, they eventually were nevertheless slaughtered by Raptor Prime, a creature of instinct and adaptation which quickly learnt to use their enemies shape-shifting abilities as well.
In the "Road to Empyre," Teddy Altman is made a mysterious offer at the cost of leaving Billy Kaplan. He accepts the offer which was to become the new ruler of the Kree-Skrull Alliance, adopting the mantle of "Dorrek VIII," and beginning the preparations of invading the Earth for "the final war."
General G'iah under the alias of "Gloria Warner" sends an encrypted message to the Skrull High Command about the progress of Project Blossom. Her mate Colonel Klrr uses the alias of "Carl Warner" and they have three daughters named Alice, Ivy, and Madison. After losing her husband to their handler and rescuing Ivy, General G'iah obtains a sample of Cotati from a lab while explaining the history of the Kree and the Skrull's feud. At their motel, they are ambushed by a Kree operative. General G'iah leads her daughters track the assailant to a Kree family. After G'iah's daughters prevent her from retaliating against them, the patriarch of the Kree family that blew up their motel room suddenly receives a message on his Kree-tech cellphone by Dorrek VIII stating to all Kree and Skrull soldiers in the field that the Kree and Skrull armadas have united to face a common enemy. They are coming to Earth to destroy them as he speaks.
Skrulls are green-skinned reptilian humanoids with large pointed ears, red or green eyes, and chins with several vertical folds in the skin below their mouths. Skrulls are known for genetic and molecular instability, and genetic diversity, due to Celestial experimentation creating the Skrull Deviants. The Skrulls are known for their physical malleability and ability to shapeshift to any size, shape, or color at will, taking on the appearance but not the characteristics of other beings and objects. Skrulls are able to assume virtually any form, be it organic (e.g., cows ) or inorganic (e.g., lamp ). As a result, the Skrulls excel at spying and infiltration. Skrulls are also able to use their shapeshifting abilities to form weapons (e.g., blades and clubs) with parts of their bodies, making them dangerous hand-to-hand combatants. Their sexual dimorphism is roughly the same of a human being with the exception that the males are mostly bald and have no facial hair, while females have full head of greenish black hair, but their sexual orientation is more complex since they can change their genders at will. Indeed, Xavin once casually said that, for a Skrull, changing gender is comparable to a human being changing their hair color. As for their reproductive system, just like the other reptilians, the females do not give live birth, but instead lay eggs and the newborns are known as "hatchlings". However, despite being reptilian, the females also possesses hair and mammary glands to nurture their newborn, just like a human being.
The Skrulls later developed the ability to render themselves undetectable when using their shapeshifting abilities, even from telepaths and those with superior senses. Reed Richards discovered a method to use technology to negate this ability and reveal a disguised Skrull's true form.
Courtesy of their advanced technology, the Skrulls have also been able to augment their abilities in certain warriors, such as the Super-Skrull and War Skrulls—an elite group who with special encoding are able to emulate the powers as well as appearance of their templates.
The Skrulls have been shown to suffer from various diseases. Some Skrulls are born with a defect that makes them unable to shapeshift, however, this defect also enhances their strength to those Skrulls including Titannus, Talos and Raze. The Skrulls can also suffer from cancer, as shown with Kl'rt and Prince Klundirk. In an alternate universe, Skyppi suffered from a form of tuberculosis which was common to the Skrull species.
Just like the Kree, the Skrulls were evolutionarily stagnant, unable to evolve. However, over the millennia Skrull mutants have been born with the K-gene as oppose to the Human Mutants X-Gene, meaning that now this might not be the case anymore.
After a Skrull dies, they revert to their original form.
The Skrull species speak a variety of languages and dialects, including: Skrullos, Skrullian, the Irdu dialect, the Tligi dialect and the Frti dialect. While the Prime Skrulls had their own language, however, it was forgotten as their species was extinct.
Since the Skrulls are a warrior race, it is hard for them to grasp almost any kind of representational art.
In Deadpool vs. Thanos #3, it is revealed that the Skrull equivalent of Hell is called Shapeless Wastes.
- Al'arok – A Skrull who posed as a Zenn-Lavian known as "Tallian Pay" to gain entrance for Skrulls to Shalla-Bal; deported after his attempt to kill the Silver Surfer failed; committed suicide to prevent capture by Kree who had disabled the Skrull starship he was on.
- Anelle – Skrull princess and the only child of Emperor Dorrek VII and Empress R´Klll; heir to the Skrull Empire and the mother of Hulkling (Dorrek VIII/Teddy Altman). Killed when Galactus consumed their Throneworld.
- Aptak – A Skrull trapped in a female Waziliah form by the Hyper-Wave Bomb. She is the lover of Nenora.
- Akilll – A Skrull representative to witness the fate of the Phoenix, on the trial by the Shi'ar.
- Ard'ran – A fourth-rank Skrull engineer who participated in the Skrull's invasion upon San Francisco. She had the telepathic abilities of any telepathic member of the X-Men
- Bag'le –
- Bartak – A Skrull who posed as Silver Surfer.
- Baryn – A Skrull who helped in liberating Prince Dezan.
- Bellok – Member of the "S-Men." Also known as Ice-Boy.
- Broodskrulls – A group of Brood/Skrull Hybrids.
- Cadre K – A group of "K-Class Deviant Skrulls" who have powers and physical appearances unlike other Skrulls. They oppose the killing of their kind by the Empire in the name of racial purity. Cadre K was led by Professor X.
- Fiz – He is a mutant Skrull who journeyed to Earth to join the X-Men. Fix assisted in the battle against Apocalypse. He led Skrull mutants to rebel against the Slave-Drivers. He left Earth with the others and Professor X to find a new homeworld. Fiz could alter his size and mass.
- Goroth – A Skrull that has stretching abilities.
- Nuro – A Skrull that has the ability to elongate his body.
- R'Tee – A Skrull that has the ability to project spikes from his arms and head.
- Spunje – A Skrull that has the ability to absorb and reflect energy.
- Z'Cann – The leader of Cadre K. She was captured by Intergalactic Council. Z'Cann is a telepath.
- Carnival – Slavers who were using a carnival to attract and capture humans as slaves. They were encountered by the New Mutants and fled in their starship, but left one behind.
- Ch'gra[volume & issue needed] – A delegate in Queen Veranke's court.
- Ch'rith – A Skrull lieutenant who participated in the Skrull's invasion upon San Francisco.
- Chrell – A Skrull Commander entrusted by Queen Veranke to train her army and slay Hulkling. Chrell possessed the powers of Mister Fantastic, Invisible Woman, Human Torch, and Thing. Chrell self-destructs trying to eliminate the Young Avengers and the Runaways.
- Criti Noll – A Skrull who was chosen to pose as Hank Pym. She tried to warn the diner owner that served her good food about the upcoming Skrull invasion only to be killed by a Skrull posing as Dum Dum Dugan and some Skrulls posing as S.H.I.E.L.D. Agents.
- Criti Noll Clone – A clone of Criti Noll who posed as Hank Pym during the Secret Invasion. She not only has the abilities matching Henry Pym, she also has the powers of Black Panther, Quicksilver and Vision.
- Dakr't –
- Dalx – Skrull diplomat and cultural attaché.
- De'Lila – A Skrull thief who sought the Inorganic Technodrone. She was then taken by the Skrulls.
- Dezan – The brother of Dorek VII. He was long imprisoned due to his pacifist nature and branded a traitor to the Skrull Empire.
- De'zean – The father of Xavin and the current Prince of Tarvax VII.
- Dharri – A Skrull lieutenant.
- Dorrek I – A Skrull King who reigned during the Skrull's first encounter with the Kree.
- Dorrek VII – Former Emperor of the Skrulls. Killed by R'Klll.
- Dro'ge – Royal Priest of Science. A Skrull scientist who experiments on the Illuminati and discovers the advancements that allowed the Empire to infiltrate Earth in a massive scale.
- Dzirot – A Skrull who participates in the Skrull invasion upon San Francisco. He dies in battle.
- Ethan Edwards – A Skrull who is a Marvel Comics pastiche of Superman.
- Evh'ser – A Skrull adjutant who participates in the Skrull invasion upon San Francisco.
- Ewe'fareek – Uncle to Xavin in the comic Runaways.
- Flaw – A Skrull who served in the Shi'ar's Death Commandos.
- Fremn –
- Fry'lu – A Skrull telepath who existed during the reign of Dorrek VII.
- Galan – The Skrull priest of science.
- Godkiller – A specially-bred female Super-Skrull that has the powers of Battleaxe, Thundra, Titania, and Volcana. She was also genetically modified to wield Beta Ray Bill's hammer Stormbreaker.
- Gorth – A Skrull who was the governor of a Skrull colony.
- Granok –
- Grrix – The Skrull Ambassador at S.W.O.R.D.
- H. Warren Craddock Imposter – A Skrull who impersonated H. Warren Craddock.
- Hagar – A Skrull who serves as High Judicator in the Skrull courts.
- Henkor –
- H'kurrek – Fanatical Skrull commander that commanded the part of the Secret Invasion directed against X-Men.
- Hokk Algol – A Skrull who is a member of the War Crimes Tribunal.
- Host –
- H'rpra – A Skrull who posed as Mockingbird.
- Hs'fld –
- Hulkling (Dorekk VIII/Teddy Altman) – Theodore "Teddy" Altman, a Kree/Skrull hybrid, the son of Anelle and the Kree Captain Mar-Vell. He is a member of the Young Avengers, with the codename "Hulkling".
- Intelligencia – A Skrull being that is a composite of the greatest Skrull minds. It was created by the Skrulls to serve as their version of the Supreme Intelligence.
- Jaketch – A Skrull who serves as an apprentice executioner of the Skrull Jury.
- Jaq –
- Jash – A Skrull lieutenant who participated in the Skrull invasion upon San Francisco.
- Jazinda – A Skrull bounty hunter and the daughter of Kl'rt the Super-Skrull.
- Jora'thrll – A Skrull who participated in the Skrull invasion upon San Francisco.
- K'eel R'kt – A royal science officer of the Skrull Empire and only survivor of the Builders destruction of the Skrull planet Hy'lt Minor. He was saved by Silver Surfer and transported to the orbital headquarters of the S.W.O.R.D. where he offered his services to them against the war with the Builders.
- K'and'rr – A Skrull commander of the 7th Fleet.
- K'arr'N –
- K'rtem – A Skrull who was an old comrade of Z'Reg.
- K'Targh –
- Knights of the Infinite – A group of Kree/Skrull hybrids who believed in a prophecy about unitying the two empires and becoming their protectors.
- Dorrek Supreme – The group's leader and the first wielder of the "Excelsior" Sword.
- M'ryn - A member of the group who founded the prophecy and father of Mur-G'nn. Killed by the wizard Moridun to be used as body for the latter.
- K'kyy – A female member of the group. She assisted in kidnapping Dorrek VIII in order for the latter to become king of the new empire.
- Mur-G'nn – A female member of the group. She assisted in kidnapping Dorrek VIII.
- Lan-Zarr – A member of the group. He guided Dorrek VIII through the test to see if he was Dorrek Supreme reincarnated.
- Varra – A female member of the group. She is killed by Moridun who wanted to possess Wiccan.
- K'vvvr – A Skrull commander who is the son of K'and'rr. He led the Skrull attack upon Wakanda.
- Kalamari – A Skrull general and ally of Paibok.
- Kalxor – A Skrull who serves as a commander for Dorekk's armada.
- Karant Kiar – A Skrull Prelate of the Skrull Empire's 7th Quadrant who was a witness at the Trial of Galactus.
- Karza –
- K'thron –
- Kerth – A Skrull who was a slave of Broker.
- Kholdsor –
- Khn'nr – A Skrull who posed as the original Captain Marvel. He turned against the other Skrulls.
- Kl'rt (Super-Skrull) – A Skrull who has the powers of the Fantastic Four and became the first Super-Skrull.
- Kly'bn – The last surviving Skrull Eternal who became a Skrull God.
- Korya – A female Skrull that is the lover of Yeti.
- Kradhal –
- Kravo –
- Kreddik –
- Krellek Council –
- Krillik –
- Krimonn the Power Prism – A Skrull transformed into a sentient power object after a failed coup d'état.
- Kylak –
- Kylor – A Skrull governor who was a competitor for the title of Skrull emperor.
- Lyja – A Skrull who infiltrated the Fantastic Four and married Johnny Storm while impersonating Alicia Masters.
- Magnitude – A Super-Skrull who has the powers of Banshee, Havok, Polaris, and Sunfire. He was sent to infiltrate the Point Men until he was exposed and killed by Delroy Garrett.
- Meg'ror –
- Melugin –
- Moloth - A Skrull who is the handler of G'iah and Klrr.
- Morfex – A Skrull who is a member of the Star Masters.
- Morrat – A Skrull warlord who is responsible for the death of Franklin Storm.
- Mrok – A Skrull soldier
- M'Lanz – A Skrull Warbride, she was tasked to stop Tel-Kar from retreating his symbiote (Venom) and getting a Skrull bio-weapon, after Tel-Kar killed her two sisters. She teamed up with Eddie Brock as Sleeper to defeat Tel-Kar.
- Muraitak – A Skrull that joined the Xandarian Nova Corps and served under Queen Adora.
- Myrn – A Skrull who is a longtime friend of Zabyk and Dezan and creator of the Hyper-Wave Bomb. After Myrn had Zabyk don an insulated armor that is said to protect him from the bomb's effects, Myrn was killed by Zabyk.
- N'ala –
- Nenora – A former Skrull spy that was trapped in Kree form after being hit by a Hyper-Wave Bomb. She is the lover of Aptak.
- Nogor – A Skrull who posed as Longshot. He is the talisman of the Skrulls and the representation of the Gods.
- Nuro –
- Orf –
- Pagon – A Skrull who posed as Elektra after Siri's demise. Killed by Echo as part of a suicide mission.
- Paibok – The Power Skrull.
- Pitt'o Nili – A Skrull who posed as the Captain America (and brainwashed to be him) who came from a downed ship in the Savage Land. Killed by Shanna the She-Devil after being hit by a poisonous dart that has a huge effect on Skrulls.
- Prime Skrull – Only surviving member of the original non-Deviant Skrull race.
- Prime Ten – Ten unnamed Skrulls who sought to obtain Mister Fantastic's matter/energy transmitter.
- Ptakr –
- Purnlr – A War Skrull who was killed in battle with the Kree.
- R'Tee –
- Rachman –
- Raksor – A Skrull who oversaw the Phoenix trial.
- Rale –
- Raze – A Skrull bounty hunter who cannot shapeshift, but he is capable with hand-to-hand combat, athletic abilities, and carries an assortment of knives and guns. He is the brother of Vranx.
- Replica – A young female Skrull who was part of the Guardians of the Galaxy.
- Ripan – A Skrull who is a second-in-command to Kylor.
- R'Klll – Wife of Dorrek VII. She killed her husband and took over in his place. She was killed when Galactus consumed Tarnak IV.
- R'Kin – A short Skrull who served as fleet mechanic. Even though he is small, he has super-strength.
- Rl'nnd – Son of Rm'twr. Rl'nnd is an undercover X-Skrull agent participating in the Secret Invasion. He has displayed the combined powers of several X-Men. Killed by Ms. Marvel.
- Rm'twr – Father of Rl'nnd. He was killed by Ms. Marvel.
- Ryga'a – Wife of Kree Soh-Larr and mother of Dorrek Supreme.
- S'Bak – A Skrull baroness.
- Samuel J. Skrull –
- Sar T'llrk – A Skrull "Soul Shepard" who participates in the Skrull invasion upon San Francisco. Killed by Cyclops.
- S'Byll – A Skrull Empress who restored the Skrull's shapeshifting ability.
- Sarnogg – A Skrull who is the son of Kl'rt.
- Sensational Hydra – A Skrull spy that works for HYDRA.
- She-Thing Imposter – An unnamed Skrull who posed as She-Thing. Killed by the Skrull Kill Krew.
- Siri – A Skrull who impersonated Elektra. She was killed by the real Elektra.
- Skragg – A Skrull pirate/mercenary that served Thanos.
- Skrull Beatles – Skrulls who posed as members of the Beatles.
- Skrull-X – A Skrullian robot who can mimic the powers of the X-Men similar to Mimic.
- Skrulls of Kral – A group of Skrulls that come from the planet Kral.
- "Skrullverine" – An unnamed Skrull working for Apocalypse who was used to replace Wolverine at the time when Apocalypse turned Wolverine into the Death. This Skrull was later killed by Death.
- Skypii – An elderly Skrull mechanic who was friends with Hercules 2300. Often assumed female form and was working as a human female model when he and Hercules first met. As the "retirement plan" for Skrull workers was execution, he went into hiding in the form of a human female. Skypii chose a statue of Hercules to be his "death-form" in eternal tribute to his friend.
- Sl'gur't – A Skrull Deviant who became a Skrull Goddess and mate to Kly'bn.
- Ssrov – A Skrull Dropshift Captain who participates in the Skrull invasion upon San Francisco.
- St'kr –
- Syrro – A Skrull major.
- Talos the Untamed – A Skrull who cannot shapeshift. However, the mutation that left him unable to shapeshift gave him enormous strength.
- Tarna – A Skrull who is part of the Agents of Cosmos. She had a symbiote partner until Venom became enraged and nearly killed it while stealing a ship, burning it with the ship's engine.
- Tenelle –
- Titannus – A Skrull who cannot shapeshift, but has been bio-engineered to possess immense strength and regeneration ability.
- Trall –
- Trl'k –
- Valth –
- Velmax (Effigy) – Founder of the super team First Line posing as a human hero Effigy with shape-shifting powers.
- Veranke – The current Skrull Queen. She posed as Spider-Woman. Killed by Norman Osborn.
- V'lrym –
- Vranx – A Skrull technology user and brother of Raze.
- Warpriest Kh'oja – A ruthless fanatic and an enemy of Power Pack.
- The Warners – A group of Skrull spies posing as an ordinary American family to make way for a future Invasion.
- G'iah - A Skrull general who operated on Earth as Gloria Warner.
- Klrr - A Skrull colonel and G'iah's wife who operated on Earth as Carl Warner. He was later killed by Moloth.
- Alkss - The middle daughter of G'iah and Klrr who operated on Earth as Alice Warner.
- Ivy Warner - The youngest daughter of G'iah and Klrr.
- Madison Warner - The oldest daughter of G'iah and Klrr.
- Warskrulls – A bunch of genetically-bred Skrulls who infiltrated the Shi'ar's culture to obtain power for the Skrull race.
- Wor'il – Delegate in Dorrek's War Council.
- Xalxor –
- Xavin – A Super-Skrull in-training and member of teenage super powered group Runaways. Xavin is the child of Prince De'zean of the Skrull planet Tarvax VII.
- X'iv – A Super-Skrull assassin sent to assassinate Hulkling. She has the powers of Daredevil, Elektra, and Cloak and Dagger.
- Yorak –
- Y'tll –
- Zabyk – A Skrull warrior who was a childhood friend of Dezan and Myrn. He had Myrn construct a Hyper-Wave Bomb to use on his enemies. Under the advice of Myrn, he donned an insulated armor (even shifting his form to fit the unusual shape of its interior) that would protect him against its effects, but it failed leaving his contorted body frozen inside and placing him in agony.
- Zankor – An alien retconned into a Skrull.
- Zedrao – A high-ranking Skrull general, notable for being on good terms with Earth's metahumans. Gave the dying Mar-Vell the highest honor in the Skrull military for being a worthy rival.
- Zirksu – A Skrull spy who assumed the alias of Diabolik.
- Zkrodd –
- Z'Reg (Crusader) – A Skrull who deserted his kind. He was formerly a member of the Initiative as the second Crusader.
- Zuhn – A Skrull who was the partner of Velmax.
- Zorr'Kiri – Skrull Goddess of Love. Killed by Gorr the God Butcher.
In Amalgam Comics, the Skrulls are the native inhabitants of Mars (having crossed DC's Green Martians with the Skrulls). Mister X (Martian Manhunter and Professor X) is an example of a Martian Skrull.
In the alternate universe limited series Earth X, Thanos dwelled in the Realm of the Dead with the entity Death. It is revealed his mother was a Skrull and Death used her secret to make him believe that Death was his mother.
Hulk: The End
In the one shot Hulk: The End, an alien robot tells an elderly Bruce Banner, the last human on the planet, that the death of the human race was widely celebrated throughout the Universe, so much so that the Skrulls and Kree had settled their differences in celebration.
The Skrullian Skymaster (real name Skymax) is a founding member of the Squadron Supreme and possesses abilities matching the Super Skrull from Earth-616. The character first appeared in flashback in Squadron Supreme #1, and appeared in more detail in Squadron Supreme: New World Order.
In an alternate reality to that in which most Marvel Comics stories take place, the Skrull Empire has been at war with the Kree Empire. In an attempt to end the bloodshed, the Skrull Empire created a means to imbue their soldiers with extraordinary powers to vanquish the Kree. Skymax was the first and the only soldier subjected to the experiments, because the World Devourer destroyed the Skrull homeworld. Sk'ym'x was in the vicinity of Earth when his starship malfunctioned. He was rescued by American astronaut Joe Ledger (who would later become Doctor Spectrum). Grateful for his rescue, Skymax gave Ledger a Power Prism which Ledger used to become the costumed adventurer Doctor Spectrum, while Skymax became the costumed adventurer known as the Skrull. Skymax and Ledger both became founding members of the Squadron Supreme, and Skymax remained with the team until member Tom Thumb repaired his ship allowing him to return to space. Skymax realized that the damage to his ship was beyond repair despite Tom Thumb's efforts, and he chose to remain on Earth in secret, searching for any possibility of other Skrull survivors. Not finding any Skymax concluded that he was indeed the last of his race; he remained on Earth.
After the Squadron Supreme returned from their adventures on Earth-616, the sorcerer Mysterium sent the surviving members to retrieve others to help in the rebellion against the Global Directorate. Doctor Spectrum was sent after "the Watcher" and discovered Skymax. Renaming himself the Skrullian Skymaster and revealing his true origin to Doctor Spectrum, Skymax rejoined the Squadron Supreme and settled with them on Utopia Isle.
The Skrulls are led by the billion-year-old Skrull Emperor Kl'rt. The Skrulls dislike being confused with the Chitauri whom they call terrorists. They have extremely advanced technology, but have not been observed to shapeshift. These Skrulls were seen only in an alternate timeline in which Reed Richards contacted their world via his teleporter; the events leading to that timeline were altered in Ultimate Fantastic Four #29 so that contact was never made.
Thirty years into the future
In an alternate timeline set thirty years into the future, the Skrull Empire was united with the Kree Empire under the rulership of Emperor Dorrek VIII and they were successful in eliminating all life on Earth.
On Earth-10021 in which the Skrulls were victorious in the end of Secret Invasion and the heroes were defeated. In this reality, the Skrulls were coexisting along with the Humans on Earth and were seen by Humans as people with good intentions. Despite helping the Humans in medical care and in technology, in reality they secretly turning the Humans into Skrulls like themselves including Norman Osborn who had married Queen Veranke. While the heroes had been hiding in Wakanda, since now they were seen as criminals. In the end, the heroes spread a virus which killed every Skrull and the Humans who had been turned into Skrulls, except Norman who had been disguised as Iron Man. Norman revealed that he had orchestrated the plan all along, but he is killed by Bucky Barnes. The heroes were then taken by the US government as world criminals.
In other media
- The Skrulls have appeared in multiple animated series based on the Fantastic Four, including the 1967 Fantastic Four series, the 1978 series, the 1994 series and the 2006 Fantastic Four: World's Greatest Heroes series.
- The Skrulls appeared in the 1990s X-Men TV series. In the episode "Mojovision" a Skrull is seen in the audience. In the episode "Dark Phoenix Saga" Lilandra consults the Skrull Empress on how to deal with Dark Phoenix's rampage.
- The Skrulls appeared in the Silver Surfer series.
- The Skrulls appeared in The Super Hero Squad Show episodes "If This Be My Thanos", "Another Order of Evil" and "Alienating with the Surfer".
- The Skrulls are featured in The Avengers: Earth's Mightiest Heroes. They are mentioned or appear in the following episodes: "459"; "Widow's Sting"; "The Man Who Stole Tomorrow"; "A Day Unlike Any Other"; "The Private War of Doctor Doom"; "Welcome to the Kree Empire"; "Who Can You Trust"; "Nightmare in Red"; "Prisoner of War"; "Infiltration"; "Secret Invasion"; "Along Came a Spider"; "Behold the Vision"; and "The Deadliest Man Alive"
- The Skrulls appear in the Avengers Assemble episode "Mojo World". Some Skrulls are seen as members of the arena audience on Mojo's spaceship.
- The Skrulls appear in the Hulk and the Agents of S.M.A.S.H. episode "Deathlok." They are led by Super-Skrull in his plot to incinerate the Earth only to be stopped by the Agents of S.M.A.S.H. and Deathlok. In the episode "Guardians of the Galaxy," the Skrulls lure the Agents of S.M.A.S.H. to another planet where they managed to capture She-Hulk, A-Bomb, and Skaar in a plot to harness their gamma energy and use it to create gamma-powered Skrull soldiers. At least two of them were created before the experiment was disrupted by Hulk, Red Hulk, Star-Lord, Rocket Raccoon, and Groot. The two gamma-powered Skrull soldiers were defeated by Red Hulk (who dove into the mixture to become large enough to fight them).
- The Skrulls appear in the film Captain Marvel. Talos the Untamed (portrayed by Ben Mendelsohn) is the main featured Skrull. In the film, one faction of the Skrulls led by Talos are portrayed as victims of a genocidal war waged by the Kree, having come to Earth to seek the aid of renegade Kree scientist Mar-Vell (here a female posing as Doctor Wendy Lawson of Project Pegasus) in devising a lightspeed engine that could take the Skrulls to safety. After Kree warrior "Vers" learns of her true identity as Carol Danvers as well as the origin of her powers coming from exposure to the energies of Mar-Vell's lightspeed engine (powered by the energies of the Tesseract) she helps defend Talos, his wife, daughter and the Skrull refugees from a Kree attack. After Yon-Rogg is defeated and sent back to the Kree Empire to carry her message to the Supreme Intelligence, Danvers leaves Earth to take the Skrulls to another part of the galaxy to find a planet for them to settle on.
- In Spider-Man: Far From Home, the post credits scene reveals that Talos and his wife Soren impersonated Nick Fury and Maria Hill during the events of the film, while the real Fury is relaxing on a Skrull spaceship.
- Actor Tye Sheridan revealed in an interview that the Skrulls were originally intended to be featured in Dark Phoenix as the main antagonists, before they were scrapped in favour of the D'Bari after the film underwent a series of reshoots.
- Skrull governor Kylor is featured as a boss in the NES Silver Surfer game. He is identified as "Emperor," although he never officially held that title in the comics.
- The Skrulls appear as enemies on their home world in Marvel: Ultimate Alliance. Skrull Deviants and Skrull Commandos are the foot soldiers that the heroes will fight. At the time when Galactus is attacking the Skrull homeworld, the heroes (who have arrived to steal the Muonic Inducer from Galactus) are forced to fight against Galactus' Punishers and Skrulls who refuse to believe that they are there to help deal with Galactus. After helping the Skrull Empress escape the planet, the player has the option of destroying the Optonic Drills that Galactus uses to drill into the planet's crust. If the player saves the Skrull homeworld from destruction, the Skrulls will form a tentative partnership with Earth and will later come to the aid of Earth when the Kree threaten to enslave humanity. Otherwise, the loss of their homeworld will destabilize the entire sector of space when the Kree and the Skrull go to war for decades costing millions of lives.
- The Skrulls appear as enemies in Fantastic Four: Rise of the Silver Surfer.
- The Skrulls appear in Marvel Super Hero Squad: The Infinity Gauntlet. Skrull Chargers and Skrull Minions serves as the foot soldiers.
- The Skrulls appear in Marvel Heroes as Chapter 10's main foe.
- The Skrulls appear in the Fantastic Four virtual pinball game for Pinball FX 2 released by Zen Studios.
- The Skrulls would have appeared in a first person Skrull-centric Avengers action game that was planned by THQ Studio Australia to coincide with the release of the first live-action movie in 2012. The game would have featured Iron Man, Captain America, the Hulk, and Thor as the main characters, with Black Widow, Hawkeye, War Machine, and Ms. Marvel appearing as unlockable characters. The story, loosely based on Secret Invasion rather than the movie, was penned by veteran comic writer Brian Michael Bendis. Though a fair amount of progress was made on the game, it was ultimately canceled after THQ Studio Australia was closed down.
- The Skrulls appear in Marvel Avengers: Battle for Earth video game.
This section needs expansion. You can help by adding to it. (October 2019)
- Schebeen, Jesse (March 6, 2019). "The Skrulls and Talos of Marvel Explained: Who Are the Captain Marvel Aliens?". IGN. Archived from the original on May 2, 2019. Retrieved July 7, 2019.
- Greg Pak and Fred van Lente (w), Rafa Sandoval (p), Roger Bonet and Greg Adams (i), Marte Gracia, Dennis Calero and Raul Trevino (col), Joe Caramagna (let), Mark Paniccia (ed). "Secret Invasion" Incredible Hercules #120 (August 2008), United States: Marvel Comics
- Glenn Herdling (w), Angel Medina (p), Bill Anderson (i), Ray Murtaugh (col), Jack Morelli (let). Blackwulf #2 (July 1994), United States: Marvel Comics
- Bill Mantlo (w), Sal Buscema (p), Ian Akin and Brian Garvey (i), Ben Sean (col), Janice Chiang (let), Bob Haras (ed). "Assistant Editor's Month 1984" Rom #50 (January 1984), United States: Marvel Comics
- Steve Englehart (w), Sal Buscema (p), Joe Staton (i), Phil Rachelson (col), Tom Orzechowski (let). Avengers #133 – 134 (March – April 1975), United States: Marvel Comics
- Scott Edelman (w), Al Milgrom (p), Terry Austin (i), Irene Vartanoff (col), Joe Rosen and Gaspar Saladino (let), Archie Goodwin (ed). Captain Marvel #51 (July 1977), United States: Marvel Comics
- Peter Gillis (w), Brian Postman (p), Kim DeMulder (i), Bob Sharen (col), Diana Albers (let), Mark Gruenwald (ed). Captain America Annual #7 (October 1983), United States: Marvel Comics
- Roy Thomas (w), Sal Buscema (p), Sam Grainger (i), Sam Rosen (let), Stan Lee (ed). "Kree-Skrull War" Avengers #89 – 97 (June 1971 – March 1972), United States: Marvel Comics
- Allan Heinberg (w), Jim Cheung (p), Dave Meikis, John Dell and Jim Cheung (i), Justin Ponsor (col), Cory Petit (let), Tom Brevoort (ed). Young Avengers #9 – 12 (December 2005 – August 2006), United States: Marvel Comics
- Roy Thomas (w), Jim Craig (p), Dave Hunt (i), George Roussos (col), John Costanza (let). Marvel Premiere #35–37 (April–August 1977), United States: Marvel Comics
- Stan Lee (w), Jack Kirby (p), George Klain and Christopher Rule (i). Fantastic Four #2 (1 January 1962), United States: Marvel Comics
- Stan Lee (w), Jack Kirby (p), Dick Ayers (i), Stan Goldberg (col), Artie Simek (let). Fantastic Four #18 (September 1963), United States: Marvel Comics
- Stan Lee (w), Jack Kirby (p), Chic Stone (i), Sam Rosen (let). Fantastic Four #32 (November 1964), United States: Marvel Comics
- Stan Lee (w), Jack Kirby (p), Chic Stone (i), Sam Rosen (let). Fantastic Four #37 (April 1965), United States: Marvel Comics
- Roy Thomas (w), Gene Colan (p), Vince Colletta (i), Arti Simek (let), Stan Lee (ed). Captain Marvel #2–3 (June–July 1968), United States: Marvel Comics
- Stan Lee (w), Jack Kirby (p), Joe Sinnott (i), Sam Rosen (let). Fantastic Four #89–93 (August 1967 – April 1970), United States: Marvel Comics
- John Byrne (w), M. D. Bright (p), Mike Gustovich (i), Petra Scotese (col), Phil Felix (let), Michael Carlin (ed). Fantastic Four Annual #18 (August 1984), United States: Marvel Comics
- Marv Wolfman (w), Keith Pollard (p), Joe Sinnott (i), Francoise Mouly (col), Jim Novak (let), Jim Shooter (ed). Fantastic Four #204 – 209 (March – August 1979), United States: Marvel Comics; Marv Wolfman (w), Carmine Infantino (p), Klaus Janson (i), Glynis Wein (col), Joe Rosen (let), Jim Shooter (ed). Nova #25 (20 February 1979), United States: Marvel Comics
- John Byrne (w), Glynis Wein (col), Jim Novak (let), Al Milgrom and Ann Nocenti (ed). Fantastic Four #257 (August 1983), United States: Marvel Comics
- Roger Stern (w), John Byrne (p), Kyle Baker (i), Christie Scheele (col), John Workman (let), Mark Gruenwald (ed). Avengers Annual #14 (30 July 1985), United States: Marvel Comics; John Byrne (w), Joe Sinnott (i), Glynis Wein (col), Jim Novak (let), Michael Carlin (ed). Fantastic Four Annual #19 (November 1985), United States: Marvel Comics
- Steve Englehart (w), Marshall Rogers and Ron Lim (p), Joe Rubinstein, Tom Christopher and Keith Williams (i), Tom Vincent (col), John Workman and Ken Bruzenak (let), Michael Higgins, Mike Rockwitz and Craig Anderson (ed). Silver Surfer v3, #1–31 (July 1987 – December 1989), United States: Marvel Comics
- Roy Thomas (w), John Buscema (p), L Joe Sinnott (i), Phil Rachelson (col), Joe Rosen (let). Fantastic Four #173–174 (August 1976), United States: Marvel Comics
- Doug Moench (w), George Perez (p), Chic Stone, Jon D'Agostino, and Mike Esposito (i), Ben Sean and George Roussos (col), Jim Novak (let), Jim Salicrup (ed). Fantastic Four Annual #15 (October 1980), United States: Marvel Comics
- John Byrne and Chris Claremont (w), Terry Austin (i), Glynis Oliver (col), Tom Orzechowski (let), Jim Salicrup and Louise Jones (ed). Uncanny X-Men #137 (10 September 1980), United States: Marvel Comics
- J.M. DeMatteis (w), Alan Kupperberg (p), Dan Green (i), Ben Sean (col), Janice Chiang (let), Jim Salicrup (ed). Avengers #209 (July 1981), United States: Marvel Comics
- Jim Stalin (w), Steve Oliff (col), Jim Novak (let), Al Milgrom (ed). "The Death of Captain Marvel" Marvel Graphic Novel #1 (12 January 1982), United States: Marvel Comics
- Mike Costa (w), Mark Bagley (p), Andrew Hennessy (i), Dono Sánchez-Almara (col), Clayton Cowles (let), Devin Lewis, Lauren Amaro and Nick Lowe (ed). Venom First Host #1–3 (August–September), United States: Marvel Comics
- Chris Claremont (w), Jim Lee (p), Scott Williams (i), Joe Rosas (col), Pat Brosseau (let), Bob Haras (ed). Uncanny X-Men #276 (May 1991), United States: Marvel Comics
- Chris Claremont (w), Jim Lee (p), Scott Williams (i), Joe Rosas (col), Tom Orzechowski (let), Bob Haras (ed). Uncanny X-Men #277 (June 1991), United States: Marvel Comics
- Tom DeFalco (w), Paul Ryan (p), Danny Bulanadi (i), Christie Scheele (col), Bill Oakley (let), Ralph Macchio (ed). Fantastic Four #358 (November 1991), United States: Marvel Comics
- Alan Davis and Terry Kavanagh (w), Alan Davis (p), Mark Farmer (i), Marie Javins (col), Richard Starkings and Saida Temofonte (let), Mark Powers (ed). "Shattering" X-Men v2, #94 (November 1999), United States: Marvel Comics
- Chris Claremont (w), Salvador Larroca (p), Tom Townsend (i), Richard Isanove (col), Richard Starkings and Comicraft (let), Mark Powers and Pete Franco (ed). "Maximum Security" Uncanny X-Men #387 (December 2000), United States: Marvel Comics
- Keith Giffen (w), Scott Kolins (p), Ariel Olivetti (i), June Chung (col), Cory Petit (let), Andy Schmidt (ed). Annihilation: Prologue #1 (May 2006), United States: Marvel Comics
- Javier Grillo-Marxuach (w), Greg Titus (p), Chris Sotomayor (col), Joe Caramagna (let), Andy Schmidt (ed). Annihilation: Super-Skrull #1 – 4 (June – September 2006), United States: Marvel Comics
- Ed Brubaker (w), Billy Tan (p), Danny Miki and Allen Martinez (i), Frank D'Armata (col), Chris Eliopoulos (let), Mike Marts, Nick Lowe and Sean Ryan (ed). "Rise and Fall of the Shi'ar Empire" Uncanny X-Men #478 (November 2006), United States: Marvel Comics
- Brian Michael Bendis (w), Leinil Yu (p), Mark Morales (i), Laura Martin (col), Chris Eliopoulos (let), Tom Brevoort (ed). Secret Invasion #1 – 8 (April – December 2008), United States: Marvel Comics
- Dan Abnett and Andy Lanning (w), Tan Eng Huat (p), Victor Olazaba (i), June Chung (col), Joe Caramagna (let), Bill Rosemann (ed). Annihilators #3 (4 May 2011), United States: Marvel Comics
- Jonathan Hickman (w), Jim Cheung (p), Mark Morales, John Livesay and David Meikis (i), Justin Ponsor (col), Chris Eliopoulos and Joe Caramagna (let), Tom Brevoort and Lauren Sankovitch (ed). Infinity #1 (14 August 2013), United States: Marvel Comics
- Jonathan Hickman (w), Jerome Opeña and Dustin Weaver (p), Justin Ponsor (col), Chris Eliopoulos (let), Tom Brevoort and Lauren Sankovitch (ed). Infinity #3 (18 September 2013), United States: Marvel Comics
- Jonathan Hickman (w), Jim Cheung (a). Infinity #6 (27 November 2013), United States: Marvel Comics
- A + X #18. Marvel Comics.
- Meet the Skrulls #1-5. Marvel Comics.
- Infinity Countdown: Darkhawk #2. Marvel Comics.
- Infinity Countdown: Darkhawk #3. Marvel Comics.
- Incoming! #1. Marvel Comics.
- Road to Empyre: The Kree/Skrull War #1. Marvel Comics.
- Roger Stern (w), John Byrne (p), Kyle Baker (i), Christie Scheele (col), John Workman (let), Mark Gruenwald (ed). Avengers Annual #14 (30 July 1985), United States: Marvel Comics
- Brian K. Vaughan (w), Takeshi Miyazawa (p), Craig Yeund (i), Christina Strain (col), Randy Gentile (let), MacKenzie Cadenhead (ed). Runaways v2, #8 (November 2005), United States: Marvel Comics
- Marvel Fact Files #2 (March 2013), United States: Marvel Comics
- Mark Gruenwald and Peter Sanderson (w), Josef Rubinstein (i), Andy Yanchus (col), Mark Gruenwald and Howard Mackie (ed). Official Handbook of the Marvel Universe v2, #12 (November 1986), United States: Marvel Comics
- Brian Michael Bendis (w), Leinil Yu (p), Dave McGaic (col), Richard Starkings and Albert Deschesne (let), Tom Brevoort (ed). "Initiative; Revolution" New Avengers #31–32 (August 2007), United States: Marvel Comics
- Brian Michael Bendis and Brian Reed (w), Jim Cheung (p), Mark Morales (i), Justin Ponsor (col), Cory Petit (let), Tom Brevoort (ed). "Secret Invasion" New Avengers: The Illuminati v2, #5 (January 2007), United States: Marvel Comics
- Brian Michael Bendis (w), Leinil Francis Yu (p), Mark Morales (i), Laura Martin, Emily Warren and Christina Strain (col), Chris Eliopoulos (let), Tom Brevoort and Jeanine Schaefer (ed). Secret Invasion #5 (10 September 2008), United States: Marvel Comics
- Chris Claremont (w), Jim Lee (p), Scott Williams (i), Joe Rosas (col), Pat Brosseau (let), Bob Harris and Suzanne Gaffney (ed). Uncanny X-Men #274 – 277 (March – June 1991), United States: Marvel Comics
- Robert Kirkman (w), Paco Medina (p), Juan Vlasco (i), Studio F (col), Virtual Calligraphy and Cory Petit (let), Tom Brevoort,Andy Schmidt,Aubrey Sitterson and Molly Lazer (ed). Marvel Team-Up v3, #12 (November 2005), United States: Marvel Comics
- Peter David (w), Roger Cruz (p), Cam Smith (i), Glynis Oliver (col), Joe Rosen (let), Bobbie Chase and James Felder (ed). Incredible Hulk #419 (July 1994), United States: Marvel Comics
- Ron Marz (w), Colleen Doran (p), Tom Christopher (i), Evelyn Stein (col), Ken Bruzenak (let), Craig Anderson (ed). Silver Surfer v3, #89 (February 1994), United States: Marvel Comics
- Ronald Byrd, Michael Hoskin, Gabriel Shechter, John Rhett Thomas, Stuart Vandal and Jeph York (w), John Rhett Thomas (ed). Skrulls! #1 (September 2008), United States: Marvel Comics
- John Byrne (w), John Byrne (p), John Byrne (i), Andy Yanchus (col), Michael Higgins (let), Dennis O'Neil and Linda Grant (ed). Alpha Flight #10 (May 1984), United States: Marvel Comics
- Dennis Hallum (w), Javier Rodriguez (p), Alvaro Lopez (i), Rachelle Rosenberg (col), Travis Lanham (let), Devin Lewis and Nick Lowe (ed). Spider-Woman v6, #3 (March 2016), Unitet States: Marvel Comics
- Bob Layton (w), Ron Lim (p), Bob Layton (i), Mike Cavallaro (col), Dave Sharpe (let), Mark Paniccia and Charlie Beckerman (ed). Hercules: Twilight of a God #1 (August 2010), United States: Marvel Comics
- Kurt Busiek (w), Jerry Ordway (p), Will Blyberg and Paul Ryan (i), Jason Wright (col), Richard Starkings and Comicraft (let), Tom Brevoort (ed). Maximum Security: Dangerous Planet #1 (October 2000), United States: Marvel Comics
- Brian Michael Bendis (w), Khoi Pham (p), Danny Miki (i), Dean White (col), Dave Lanphear (let), Tom Brevoort (ed). Mighty Avengers #16 (September 2008), United States: Marvel Comics
- Runaways Vol. 2 #8. Marvel Comics.
- Fantastic Four Annual #18. Marvel Comics.
- New Avengers #40. Marvel Comics.
- What If...? World War Hulk #1. Marvel Comics.
- Mighty Avengers #12. Marvel Comics.
- Incredible Hercules #120. Marvel Comics.
- Blackwulf #10. Marvel Comics.
- U.S.Avengers #12. Marvel Comics.
- Secret Invasion Who Do You Trust? #1. Marvel Comics.
- Incredible Hercules #117. Marvel Comics.
- Thor: God of Thunder #3. Marvel Comics.
- Deadpool vs. Thanos #3. Marvel Comics.
- Silver Surfer vol. 3 #2. Marvel Comics.
- Silver Surfer vol. 3 #3
- Fantastic Four #37. Marvel Comics.
- Fantastic Four #257
- Silver Surfer vol. 3 #6. Marvel Comics.
- Uncanny X-Men #137. Marvel Comics.
- Secret Invasion: X-Men #4. Marvel Comics.
- Fantastic Four #347. Marvel Comics.
- Avengers Annual #14. Marvel Comics.
- X-Men: First Class #6. Marvel Comics.
- X-Men Vol 4 #21. Marvel Comics.
- X-Men vol. 2 #97. Marvel Comics.
- X-Men vol. 2 #95. Marvel Comics.
- Uncanny X-Men #387. Marvel Comics.
- Maximum Security: Dangerous Planet #1. Marvel Comics.
- New Mutants #92. Marvel Comics.
- Secret Invasion: X-Men #2. Marvel Comics.
- Secret Invasion: Runaways/Young Avengers #1–3. Marvel Comics.
- Mighty Avengers #15. Marvel Comics.
- Secret Invasion #1. Marvel Comics.
- Fantastic Four #347. Marvel Comics.
- FF Fifty Fantastic Years #1. Marvel Comics.
- Spider-Man / Fantastic Four #3. Marvel Comics.
- Avengers Annual #14. Marvel Comics.
- Avengers: The Initiative #18. Marvel Comics.
- Avengers #133. Marvel Comics.
- Fantastic Four #18. Marvel Comics.
- Secret Invasion: X-Men #1. Marvel Comics.
- Marvel Knights: Spider-Man #13. Marvel Comics.
- Uncanny X-Men #467. Marvel Comics.
- New Avengers #44. Marvel Comics.
- Secret Invasion: Thor #2. Marvel Comics.
- Avengers #259. Marvel Comics.
- X-Men Vol. 2 #95. Marvel Comics.
- Avengers #42. Marvel Comics.
- Fantastic Four #206. Marvel Comics.
- Fantastic Four #249. Marvel Comics.
- Secret Invasion: X-Men #1–4. Marvel Comics.
- Force Works #13. Marvel Comics.
- Uncanny X-Men #269. Marvel Comics.
- New Avengers: The Reunion #2. Marvel Comics.
- Ms. Marvel Vol. 2 #25. Marvel Comics.
- Young Avengers #1. Marvel Comics.
- Captain Marvel vol. 5 #8. Marvel Comics.
- Fantastic Four #206. Marvel Comics.
- Secret Invasion: X-Men #2–4. Marvel Comics.
- She-Hulk Vol. 2 #22. Marvel Comics.
- Infinity: Against the Tide: Infinite Comic #1-2. Marvel Comics.
- Black Panther Vol. 4 #39. Marvel Comics.
- Avengers: The Initiative #15. Marvel Comics.
- Fantastic Four Annual #24. Marvel Comics.
- New Avengers Vol. 4 #3. Marvel Comics.
- Black Panther vol. 4 #41. Marvel Comics.
- Fantastic Four #358. Marvel Comics.
- Avengers #96. Marvel Comics.
- Fantastic Four #262. Marvel Comics.
- A + X #13. Marvel Comics.
- Force Works #15. Marvel Comics.
- Avengers Spotlight #25. Marvel Comics.
- Civil War: The Return #1. Marvel Comics.
- Runaways Vol. 2 #14. Marvel Comics.
- Marvel: The Lost Generation #12. Marvel Comics.
- Force Works #14. Marvel Comics.
- She-Hulk Vol. 2 #3. Marvel Comics.
- X-Men First Class #6. Marvel Comics.
- Avengers #69. Marvel Comics.
- Fantastic Four #265. Marvel Comics.
- Avengers: The Initiative #14. Marvel Comics.
- Meet the Skrulls #1. Marvel Comics.
- Star Masters #1
- Fantastic Four #32. Marvel Comics.
- Deadpool #24. Marvel Comics.
- Venom First Host. Marvel Comics.
- Nova vol. 2 #15. Marvel Comics.
- Annihilation: The Nova Corps Files #1. Marvel Comics.
- X-Factor vol. 5 #33. Marvel Comics.
- New Avengers #29. Marvel Comics.
- Secret Invasion #1. Marvel Comics.
- Blackwulf #2. Marvel Comics.
- Silver Surfer Vol. 3 #2. Marvel Comics.
- X-Men Vol. 2 #97. Marvel Comics.
- Lunatik #2. Marvel Comics.
- X-Men #37. Marvel Comics.
- Silver Surfer vol. 3 #89. Marvel Comics.
- Guardians of the Galaxy #9. Marvel Comics.
- Silver Surfer vol. 3 #4. Marvel Comics.
- Fantastic Four #206. Marvel Comics.
- New Avengers Vol 4 #4. Marvel Comics.
- Annihilation: Super-Skrull #1. Marvel Comics.
- Silver Surfer vol. 3 #13. Marvel Comics.
- Captain America vol. 3 #3. Marvel Comics.
- Captain Marvel vol. 1 #25. Marvel Comics.
- Wisdom #6. Marvel Comics.
- Captain Britain and MI-13 #4. Marvel Comics.
- Super Spy Weekend: John the Skrull, Comic Book Resources, March 8, 2008
- Fantastic Four #204. Marvel Comics.
- Fantastic Four #90. Marvel Comics.
- Uncanny X-Men #371. Marvel Comics.
- Astonishing X-Men Vol. 2 #3. Marvel Comics.
- Hercules Vol. 2 #2. Marvel Comics.
- Hercules: Twilight of a God #4. Marvel Comics.
- Fantastic Four Annual #24. Marvel Comics.
- Silver Surfer Vol. 3 #27. Marvel Comics.
- Incredible Hulk vol. 2 #418. Marvel Comics.
- Venom Space Knight #7. Marvel Comics.
- Namor the Sub-Mariner #18. Marvel Comics.
- Marvel Team-Up Vol. 3 #2. Marvel Comics.
- Marvel: The Lost Generation #12. Marvel Comics.
- New Avengers #1. Marvel Comics.
- Avengers: The Initiative #16. Marvel Comics.
- Skrulls Vs. Power Pack #1. Marvel Comics.
- Uncanny X-Men #275
- Fantastic Four #209. Marvel Comics.
- Runaways Vol. 2 #7. Marvel Comics.
- Secret Invasion: Runaways/Young Avengers #1. Marvel Comics.
- Silver Surfer Vol. 3 #6. Marvel Comics.
- Silver Surfer Vol. 3 #28. Marvel Comics.
- Amazing Adult Fantasy #7. Marvel Comics.
- Marvel: The Lost Generation #11. Marvel Comics.
- Marvel Graphic Novel #1. Marvel Comics.
- Marvel Premiere #35
- Marvel Team-Up Vol. 3 #20. Marvel Comics.
- Marvel: The Lost Generation #4. Marvel Comics.
- Mark Waid and Gerard Jonespenciller=Howard Porter (w), John Dell (i), Gloria Vasquez and Heroic Age Studios (col), Chris Eliopoulos (let), Ruben Diaz, Brian Augustyn and Ali Morales (ed). JLX #1 (April 1996), United States: Marvel Comics
- Jim Krueger and Alex Ross (w), John Paul Leon (p), Bill Reinhold (i), Matt Hollingsworth (col), Marie Javins and Polly Watson (ed). Earth X #0–12, X (Mar. 1999 – June 2000), United States: Marvel Comics
- Jim Krueger and Alex Ross (w), Doug Braithwaite (p), Bill Reinhold (i), Peter Pantazis and Benjamin Garcia (col), Todd Klein (let), Mike Marts (ed). Universe X X (November 2001), United States: Marvel Comics
- Peter David (w), Dale Keown (p), Joe Weems and Livesay (i), Avalon Studios and Dan Kemp (col), John Workman (let), Bobbie Chase, Tom Brevoort and Marc Sumerak (ed). Incredible Hulk: The End #1 (August 2002), United States: Marvel Comics
- Len Kaminski (w), Anthony Williams (p), Andy Lanning (i), Tom Smith (col), Richard Starkings and Comicraft (let), Tom Brevoort (ed). Squadron Supreme: New World Order (1998), United States: Marvel Comics
- Squadron Supreme #1. Marvel Comics.
- Squadron Supreme: New World Order (September 1998). Marvel Comics.
- Mark Millar (w), Greg Land (p), Matt Ryan (i), Justin Ponsor (col), Randy Gentile (let), Ralph Macchio, John Barber and Nicole Wiley Boose (ed). Ultimate Fantastic Four #27 – 29 (April–June 2004), United States: Marvel Comics
- DeFalco, Tom (w), Ryan, Paul (p), Milgrom, Al (i). "Wrecked and Walloped by the Wizard's Warriors" Fantastic Five 2 (November 1999)
- Kieron Gillen (w), Joe Bennett (p), Scott Hanna (i), Guru-eFX (col), Joe Caramagna (let), Mark Paniccia (ed). Iron Man v5, #18 (November 2013), United States: Marvel Comics
- Kevin Grevioux and Karl Bollers (w), Pow Rodrix (p), Frank Martin (col), Jeff Powell (let), Justin Gabriel and A.J. Fierro (ed). "Secret Invasion" What If? (3 December 2009), United States: Marvel Comics
- Walker, Glenn (March 9, 2014). "Avengers Assemble S01 E18: Mojo World - Review". Biff Bam Pop. Archived from the original on August 10, 2015. Retrieved July 7, 2019.
- Lang, Brent (July 22, 2017). "'Captain Marvel' Will Be Set in the '90s With Skrulls as Villains". Variety. Archived from the original on July 23, 2017. Retrieved July 22, 2017.
- Coggan, Devan (September 5, 2018). "See 10 Exclusive Images From Captain Marvel". Entertainment Weekly. Retrieved September 5, 2018.
- Robinson, Joanna (July 2, 2019). "That Spider-Man: Far From Home End of Credits Reveal, Explained". Vanity Fair. Archived from the original on July 2, 2019. Retrieved July 2, 2019.
- "Original Dark Phoenix Ending Included Skrulls And A 'Huge' New York Battle, According To Tye Sheridan". CINEMABLEND. June 10, 2019.
- Vertura, Travis (April 25, 2017). "Bad Games with Good Music: Part 1". Geeks WorldWide. Archived from the original on July 5, 2017. Retrieved July 7, 2019.
- Fahey, Mike (April 29, 2011). "Marvel Pinball Celebrates 50 Years of the Fantastic Four". Kotaku. Archived from the original on July 6, 2019. Retrieved July 6, 2019.
- Dornbush, Jonathan (November 30, 2015). "See the history of the canceled Avengers video game". Entertainment Weekly. Archived from the original on April 6, 2016. Retrieved July 6, 2019.
- Skipper, Ben (November 30, 2015). "Marvel's Avengers: Cancelled first person game revealed with concept footage". International Business Times. Archived from the original on March 4, 2016. Retrieved July 6, 2019.
- "Marvel Avengers: Battle for Earth Q&A". Wired. January 29, 2013. Archived from the original on December 22, 2016. Retrieved July 6, 2019.
- WatchMojo. "Top 10 alien races in comics".