Southern right whale
|Southern right whale|
The southern right whale (Eubalaena australis) is a baleen whale, one of three species classified as right whales belonging to the genus Eubalaena. Like other right whales, the southern right whale is readily distinguished from others by the callosities on its head, a broad back without a dorsal fin, and a long arching mouth that begins above the eye. Its skin is very dark grey or black, occasionally with some white patches on the belly. The right whale's callosities appear white due to large colonies of cyamids (whale lice). It is almost indistinguishable from the closely related North Atlantic and the North Pacific right whales, displaying only minor skull differences. It may have fewer callosities on its head and more on its lower lips than the two northern species. Approximately 10,000 southern right whales are spread throughout the southern part of the Southern Hemisphere.
The size of an adult female is 15 m (49 ft) and can weigh up to 47 tonnes (46 long tons; 52 short tons), with the larger records of 17.5–18 m (57–59 ft) in length and 80 tonnes (79 long tons; 88 short tons) in weight, making them slightly smaller than other right whales in Northern Hemisphere. The testicles of right whales are likely to be the largest of any animal, each weighing around 500 kg (1,100 lb). This suggests that sperm competition is important in the mating process. Right whales cannot cross the warm equatorial waters to connect with the other (sub)species and (inter)breed: their thick layers of insulating blubber make it impossible for them to dissipate their internal body heat in tropical waters. However, based on historical records and unconfirmed sightings in modern periods, E. australis actually do occur in equatorial waters.
The proportion and numbers of molten-coloured individuals are notable in this species compared with the other species in the Northern Hemisphere. Some whales remain white even after growing up.
- 1 Taxonomy
- 2 Behavior
- 3 Population and distribution
- 4 Whaling
- 5 Conservation
- 6 Whale watching
- 7 See also
- 8 Gallery
- 9 Footnotes
- 10 References
- 11 External links
The right whales were first classified in the Balaena genus in 1758 by Carl Linnaeus, who at the time considered all of the right whales (including the bowhead) as a single species. Through the 1800s and 1900s, in fact, the Balaenidae family has been the subject of great taxonometric debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. In the early whaling days, they were all thought to be a single species, Balaena mysticetus.
The southern right whale was initially described as Balaena australis by Desmoulins in 1822. Eventually, it was recognized that bowheads and right whales were in fact different, and John Edward Gray proposed the Eubalaena genus for the right whale in 1864. Later, morphological factors such as differences in the skull shape of northern and southern right whales indicated at least two species of right whale—one in the Northern Hemisphere, the other in the Southern Ocean. As recently as 1998, Rice, in his comprehensive and otherwise authoritative classification, Marine mammals of the world: systematics and distribution, listed just two species: Balaena glacialis (all of the right whales) and Balaena mysticetus (the bowheads).
In 2000, Rosenbaum et al. disagreed, based on data from their genetic study of DNA samples from each of the whale populations. Genetic evidence now clearly demonstrates that the northern and southern populations of right whale have not interbred for between 3 million and 12 million years, confirming the southern right whale as a distinct species. The northern Pacific and Atlantic populations are also distinct, with the North Pacific right whale being more closely related to the southern right whale than to the North Atlantic right whale.
It is believed that the right whale populations first split because of the joining of North and South America. The rising temperatures at the equator then created a second split, into the northern and southern groups, preventing them from interbreeding.
The cladogram is a tool for visualizing and comparing the evolutionary relationships between taxa. The point where a node branches off is analogous to an evolutionary branching – the diagram can be read left-to-right, much like a timeline. The following cladogram of the Balaenidae family serves to illustrate the current scientific consensus as to the relationships between the southern right whale and the other members of its family.
Other junior synonyms for E. australis have included B. antarctica (Lesson, 1828), B. antipodarum (Gray, 1843), Hunterus temminckii (Gray, 1864), and E. glacialis australis (Tomilin, 1962) (see side panel for more synonyms).
Like other right whales, they are rather active on the water surface, and being curious towards human vessels. Southern rights appear to be more active and tend to interact with humans more than the other two northern species. One behavior unique to the southern right whale, known as sailing, is that of using their elevated flukes to catch the wind, remaining in the same position for considerable amount of time. It appears to be a form of play and is most commonly seen off the coast of Argentina and South Africa. Some other species such as humpback whales are also known to display. Right whales are often seen interacting with other cetaceans, especially humpback whales and dolphins. There is a record of a southern right and a humpback thought to be involved in mating activities off Mozambique.
Population and distribution
The southern right whale spends summer in the far Southern Ocean feeding, probably close to Antarctica. Feeding can be occurred if there are any chances even in temperate waters such as along Buenos Aires. It migrates north in winter for breeding and can be seen by the coasts of Argentina, Australia, Brazil, Chile, Namibia, Mozambique, Peru, Tristan de Cunha, Uruguay, Madagascar, New Zealand and South Africa. The total population is estimated to be around 10,000. Since hunting ceased, stocks are estimated to have grown by 7% a year. It appears that the South American, South African and Australasian groups intermix very little if at all, because maternal fidelity to feeding and calving habitats is very strong. The mother also passes these choices to her calves. If the later mentioned sighting was truly of E. australis, this species could cross the Equator on irregular occaions.
The most recent population estimates, published by National Geographic in October 2008, put the southern whale population at 10,000. The estimate of 7,000 followed a March 1998 IWC workshop. Researchers used data about adult female populations from three surveys (one in each of Argentina, South Africa and Australia, collected during the 1990s) and extrapolated to include unsurveyed areas, number of males and calves using available male:female and adult:calf ratios to give an estimated 1999 figure of 7,500 animals.
Many locations throughout Southern Hemisphere were named after presences of southern rights and associations with mankind where whales are rare today in some of these locations, such as Walvis Bay, Punta Ballena, Right Whale Bay, Otago Harbour, Whangarei Harbour, South Taranaki Bight, Wineglass Bay, and so on.
Hermanus in South Africa has become known as a mecca for whale watching, during the Southern Hemisphere winter months (June - October) the southern right whales migrate to the coastal waters of South Africa, with in excess of 100 whales known to be in the Hermanus area. Whilst in the area, the whales can be seen with their young as they come to Walker Bay to calve and mate. Many behaviours such as breaching, sailing, lobtailing, or spyhopping can be witnessed. In False Bay whales can be seen from the shore from July to October while both Plettenberg Bay and Algoa Bay are also home to the southern right whales from July to December. They can be viewed from land as well as by boat with licensed operators conducting ocean safaris throughout the year.
Recent increases in numbers of whales visiting the north-eastern part of South Africa, the so-called Dolphin Coast such as around Ballito and off Umdloti Beach, indicates the whales' normal ranges are expanding and that re-colonising historical habitats will likely continue as more whales migrate further north.
Namibia and Angola
In Namibia, most of confirmed whales are restricted to the south of Luderitz, the southern edge of the country, and only a handful animals, but with good increases in numbers, venture further north of historical breeding grounds such as at Walvis Bay. Until the cease of any sorts of hunting including illegal mass hunts by Soviet Union, whales were rare along Namibian shores as there had been no sighting records north of Orange River until 1971, and the first of calving activities was confirmed as late as the 1980s.
Madagascar and Mozambique
In contrast to the case in South Africa, right whales are becoming more common migrants but within very small numbers along Mozambique and Madagascar. Whales were historically seen in large numbers at various locations such as off Durban, in Delagoa/Maputo Bay and Inhaca Island, and around the Bazaruto Archipelago. The first sighting off Mozambique since the end of whaling was in 1997.
In Brazil, more than 300 individuals have been cataloged through photo identification (using head callosities) by the Brazilian Right Whale Project, maintained jointly by Petrobras (the Brazilian state-owned oil company) and the International Wildlife Coalition. The State of Santa Catarina hosts a concentration of breeding and calving right whales from June to November, and females from this population also calve off Argentinian Patagonia and Uruguay. In recent years, possibly due to changing habitat environments by human activities and conflicts with local fisheries, the number of whales visiting the coasts is decreasing.
During the 2012 annual meeting of the International Whaling Commission's Scientific Committee, data was presented regarding the continued phenomena of southern right whale strandings and high rate of mortality at Península Valdés, Argentina. Between 2003 and 2011, a total of 482 dead right whales were recorded at Península Valdés. There were at least 55 whale deaths in 2010, and 61 in 2011. As in previous years, the vast majority of strandings were calves of the season. There are increasing sightings along various other locations in recent years such as on Golfo San Jorge, Tierra del Fuego, Puerto Deseado, Mar del Plata, Miramar, Buenos Aires, Bahía Blanca, and so on.
In Uruguay, coastal areas such as Punta del Este host congregating sites for whales in breeding seasons, but not likely as calving grounds. Their recovery helped create a whale-sanctuary off Latin America; the creation of this protected area had been prevented for nearly a decade by pro-whaling nations such as Japan.
Chile and Peru
For the critically endangered Chile/Peru population, the Cetacean Conservation Center (CCC) has been working on a separate program for right whales. This population, containing no more than 50 or less individuals, is under threats of increasing ship lanes and fishing industries. Aside from vagrants' records, Peru's coastlines possibly host one of the northernmost confirmed range of the species along with Gabon, Senegal, Tanzania, Brazilian coasts, Indian Ocean, and tropical waters including South Pacific Islands. The Alfaguara project targeting cetaceans in Chiloe may possibly target this species as well in the future. Forging grounds of this population is currently undetected, but possibly down south of Caleta Zorra to southern fiords such as from Penas Gulf to Beagle Channel. Some hope arising for establishment of new tourism industry in eastern side of the Strait of Magellan most notably vicinity to Cape Virgenes as the number of sightings increases. It is unknown whether these increases are due to re-colonisation by whales from the Patagonian population. Whales are still rare in the Beagle Channel as well.
Historically, populations in Oceanian regions had been very robust. There were stories of early settlers complaining that sounds of cavorting whales kept them awake at night in various locations such as on Wellington Bay and River Derwent. Satellite tracking conducted suggests that there are at least some interactions between populations in these two nations, but the extent thereof is unknown.
Southern right whales can be found in many parts of southern Australia, where the largest population is found at the Head of the Bight in South Australia, a sparsely populated area in the middle of the Nullarbor Plain. Over 100 individuals are seen there annually from June to October. Visitors can view the whales from cliff-top boardwalks and lookouts, with whales swimming almost directly below. A more accessible South Australian location for viewing whales is Encounter Bay where the whales can be seen just off the beaches of the Fleurieu Peninsula, centred around the surfing town of Middleton. A newer nursery ground has been established on Eyre Peninsula, especially at Fowlers Bay. Numbers are much smaller at these locations compared to the Bight, with an average of a couple of whales per day, but in recent years there have been regular sightings of more than ten whales at a time off Basham Beach, near Middleton. The South Australian Whale Centre at Victor Harbor has information on the history of whaling and whale watching in the area, and maintains an on-line database of whale sightings. Whale numbers are scarcer in Victoria where the only established breeding ground which whales use each year, in very small numbers, is at Warrnambool. However, as the whales do seem to be increasing in number generally, but not showing any dramatic increases at Warrnambool, they may be extending their wintering habitats into other areas of Victoria, where the numbers of sightings are slowly increasing. These areas include around Melbourne, such as in Port Phillip Bay, along Waratah Bay, at Ocean Grove, Warrnambool, on Mornington Peninsula, in Apollo Bay, and on Gippsland coasts and at Wilsons Promontory. Tasmania is another, newer, wintering ground showing dramatic increases in recent years. The waters off the Western Australia, New South Wales, and Queensland coasts had previously been inhabited by whales. Their historical range was much wider and was spread around the southern coast of the continent, extending up to Exmouth and Shark Bay on the west coast, and to Hervey Bay and Moreton Bay on the east coast. The east coast population is still endangered and very small (in low-tens), contributing in small numbers and limited re-colonization, but increases have been confirmed in many areas such as the vicinity of Sydney Harbor, Port Stephens, Twofold Bay, Jervis Bay, Broulee, Moruya River, Narooma, Byron Bay, and so on, and there have been 12 foraging areas officially announced.[not in citation given]
Some whales can be seen around in sub-Antarctic regions such as Macquarie Island. It is unknown whether historical oceanic habitats such as Norfolk Island and Lord Howe Island (known as the "Middle Ground" for whalers) would be re-colonised by Australian populations in the future.
Many features are still unknown about right whale populations in New Zealand waters. However, studies by the Department of Conservation and sightings reported by locals helped to deepen understanding. Scientists used to believe there was a very small, remnant population of southern right whales inhabiting New Zealand's main islands (North and South Island), containing probably 11 reproductive females. In winter, whales migrate north to New Zealand waters and large concentrations occasionally visit the southern coasts of South Island. Bay areas along Foveaux Strait from Fiordland region to northern Otago are important breeding habitats for right whales, especially Preservation Inlet, Te Waewae Bay, and Otago Peninsula. Calving activities are observed all around the nation, but with more regularity around North Island shores from the Taranaki coast in the west to Hawke's Bay, Bay of Plenty in the east, and areas in Hauraki Gulf such as Firth of Thames or Bay of Islands in the north. The population at the sub-Antarctic Auckland Islands is showing a remarkable recovery while the recovery state in Campbell Islands is slower. There are arious parts of the nation where large numbers of whales were seen historically, but sightings are less common nowadays. These areas include the Marlborough Region especially from Clifford Bay and Cloudy Bays to Port Underwood, Golden Bay, Awaroa Bay, and coastlines on West Coast and Hokianga Harbour in Northland. Other than a handful of confirmed observations, very little information are available for whether whales still migrating to historical oceanic habitats of Kermadec Islands and Chatham Islands.
Recent study revealed that the right whale populations from New Zealand's main islands and the sub-Antarctic islands interbreed, though it is still unknown whether the two stock originally came from a single population.
In oceanic islands and offshore waters other than the abovementioned areas, very little about the presence and recovery status of southern right whales is known. Whales' historical ranges were much broader than today as whales were known to occur at lower latitude areas such as around Pacific Islands during whaling days, and were also frequent pelagic waters at lower latitude of central Indial Ocean near Equator line, around the area locates at middle between the atolls and islands in west (Diego Garcia, Egmont Islands, and Great Chagos Bank) and Cocos Island in southeast, being comparable to the range of the Chile/Peru stock, the northernmost of all the populations known today. Whales were also taken off Gabon, Kiribati, and Tanzania, and there have been a few of unconfirmed sightings in recent years including one by Jim Darling, a renowned whale researcher. Due to illegal whaling by the Soviet Union, the recovery of many stocks including the population off Tristan da Cunha and adjacent areas such as Gough Island had been severely hindered, resulting in relatively few numbers of visiting animals. Whales' numbers visiting historical sub-Antarctic Islands show drastic differences in quantity for respective locations: well-recovering at New Zealand Subantarctic Islands while less successful at Macquarie Island, South Georgia and Falkland Islands, and likely to be slower recovery-rates at other archipelagos such as Crozet Islands and Marion Islands.
By 1750 the North Atlantic right whale was as good as extinct for commercial purposes and the Yankee whalers moved into the South Atlantic before the end of the 18th century. The southernmost Brazilian whaling station was established in 1796, in Imbituba. Over the next one hundred years, Yankee whaling spread into the Southern and Pacific Oceans, where the American fleet was joined by fleets from several European nations.
The southern right whale had been coming to New Zealand waters in large numbers before the 19th century, but was extensively hunted from 1830-1850. Hunting gradually declined with the whale population and then all but ended in coastal New Zealand waters. The beginning of the 20th century brought industrial whaling, and the catch grew rapidly. By 1937, according to whalers' records, 38,000 were captured in the South Atlantic, 39,000 in the South Pacific, and 1,300 in the Indian Ocean. Given the incompleteness of these records, the total take was somewhat higher.
As it became clear that stocks were nearly depleted, right whaling was banned in 1937. The ban was largely successful, although some illegal whaling continued for several decades. Madeira took its last two right whales in 1968. Illegal whaling continued off the coast of Brazil for many years and the Imbituba station processed right whales until 1973. The Soviet Union admitted illegally taking over 3,300 during the 1950s and 1960s, although it only reported taking 4. It was also revealed that Japan was supporting these destructive hunts by neglecting and disregarding monitoring obligations. Furthermore, there were agreements between Japan and the Soviet Union to keep their illegal mass whaling activities in foreign/international protected waters in confidence.
Whales began to be seen again in Australian and New Zealand waters from the early 1960s. It is claimed that if the illegal hunts by the Soviet Union had never happened, the New Zealand population would be three or four times larger than its current size.
The southern right whale, listed as "endangered" by CITES, is protected by all countries with known breeding populations (Argentina, Australia, Brazil, Chile, New Zealand, South Africa and Uruguay). In Brazil, a federal Environmental Protection Area encompassing some 1,560 km2 (600 sq mi) and 130 km (81 mi) of coastline in Santa Catarina State was established in 2000 to protect the species' main breeding grounds in Brazil and promote regulated whale watching.
The southern right whale is listed on Appendix I of the Convention on the Conservation of Migratory Species of Wild Animals (CMS) as this species has been categorized as being in danger of extinction throughout all or a significant proportion of their range.
One possibly significant contributor to the calf mortality rate has alarmed scientists – since at least 1996, kelp gulls off the coast of Patagonia have been observed attacking and feeding on live right whales. The kelp gull uses its powerful beak to peck down several centimetres into the skin and blubber, often leaving the whales with large open sores – some of which have been observed to be half a meter in diameter. This predatory behavior, primarily targeted towards mother/calf pairs, has been continually documented in Argentinian waters, and continues today. Observers note that the whales are spending up to a third of their time and energy performing evasive maneuvers – therefore, mothers spend less time nursing, and the calves are thinner and weaker as a result. Researchers speculate that many years ago, waste from fish processing plants allowed the gull populations to soar. Their resulting overpopulation, combined with reduced waste output, caused the gulls to seek out this alternative food source. Scientists fear that the gulls' learned behaviour could proliferate, and the IWC Scientific Committee has urged Brazil to consider taking immediate action if and when similar gull behaviour is observed in their waters. Such action may include the removal of attacking gulls, following Argentina's lead in attempting to reverse the trend.
The southern right whale has made Hermanus, South Africa one of the world centers for whale watching. During the winter months (June to October), southern right whales come so close to the shoreline that visitors can watch them from the shore as well as from strategically placed hotels. Hermanus also has two boat–based whale watching operators. The town employs a "whale crier" (cf. town crier) to walk through the town announcing where whales have been seen. Southern right whales can also be watched at other winter breeding grounds. In False Bay whale-watching can be done from the shore or from the boats of licensed operators in Simon's Town. Plettenberg Bay along the Garden Route of South Africa is another mecca for whale watching not only for southern rights (July to December)but throughout the year. There are both land based and ocean safaris boat based whale encounters on offer in this beautiful town. Southern right whales can also be seen off the coast of Port Elizabeth with marine eco tours running from the Port Elizabeth harbour, as some southern right whales make Algoa Bay their home for the winter months.
Whales are occasionally observed during tours in Namibia, Mozambique and Madagascar, where sighting rates along Namibian coasts shows dramatic increases in the recent years.
In Brazil, Imbituba in Santa Catarina has been recognized as the National Right Whale Capital and holds annual Right Whale Week celebrations in September, when mothers and calves are more often seen. The old whaling station there is now a museum that documents the history of right whales in Brazil. In Argentina, Península Valdés in Patagonia hosts (in winter) the largest breeding population, with more than 2,000 catalogued by the Whale Conservation Institute and Ocean Alliance. As in the south of Argentina, the whales come within 200 m (660 ft) of the main beach in the city of Puerto Madryn and form a part of the large ecotourism industry. Uruguay's Parliament on 4 September 2013, has become the first country in the world to make all of its territorial waters a safehaven for whales and dolphins. Every year, dozens of whales are sighted, especially in the departments of Maldonado and Rocha during the months of winter. Swimming activities for commercial objectives had been banned in the area in 1985, but is legalized in Gulf of San Matías where this is the only location in the world for tourists to be permissioned to swim with the species. Land-based watching and occasional kayaking with whales activities are Various other locations where is not renown for whale-watching industries as much as Puerto Madryn and with less restrictions on approaching whales, such as at Puerto Deseado, Mar del Plata, and Miramar in Buenos Aires
Though their number is dangerously small, land-based sightings of whales are on increase in recent years off Chile and Peru, with some hope to create new tourism industries, especially in the strait of Magellan most notably around Cape Virgenes.
In Australia's winter and spring, southern right whales can be seen from the Bunda Cliffs and Twin Rocks, both along the remote Great Australian Bight in South Australia. In Warrnambool, Victoria, there exists a nursery which is popular with tourists in the winter and spring. Their normal range is extending as the species is recovering and re-colonizing to other areas of the continent, especially around coastal waters of New South Wales and Tasmania. In Tasmania, the first birth record since the 19th century and several more following births were recorded in River Derwent since after in 2010.
For same reason, southern rights may provide chances for public to observe whales from shore on New Zealand coasts with more regularity than in the past decades, especially in southern Fiordland, Southland to Otago coast, and on North Island coast especially in Northland and some other locations such as Bay of Plenty and South Taranaki Bight. Births of calves could have always been occurred on main islands' coast, but firstly confirmed of two cow-calf pairs in 2012.
In the Subantarctic Islands and general vicinity to Antarctica, where few regulations exist or are enforced, whales can be observed on expedition tours with increasing probabilities. The Auckland Islands are a specially designated sanctuary for right whales, where any kind of whale-watching tourism is prohibited without permission.
Eubalaena australis on Wikimedia Commons.
- The use of Order Cetartiodactyla, instead of Cetacea with Suborder Mysticeti, is favored by most evolutionary mammalogists working with molecular data and is supported the IUCN Cetacean Specialist Group and by Taxonomy Committee of the Society for Marine Mammalogy, the largest international association of marine mammal scientists in the world. See Cetartiodactyla and Marine mammal articles for further discussion.
- Mead, J.G.; Brownell, R.L., Jr. (2005). "Order Cetacea". In Wilson, D.E.; Reeder, D.M. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 723–743. ISBN 978-0-8018-8221-0. OCLC 62265494.
- Reilly, S.B., Bannister, J.L., Best, P.B., Brown, M., Brownell Jr., R.L., Butterworth, D.S., Clapham, P.J., Cooke, J., Donovan, G.P., Urbán, J. & Zerbini, A.N. (2008). "Eubalaena australis". IUCN Red List of Threatened Species. Version 2009.1. International Union for Conservation of Nature. Retrieved 12 September 2009.
- Agnarsson, I.; May-Collado, LJ. (2008). "The phylogeny of Cetartiodactyla: the importance of dense taxon sampling, missing data, and the remarkable promise of cytochrome b to provide reliable species-level phylogenies". Mol Phylogenet Evol. 48 (3): 964–985. doi:10.1016/j.ympev.2008.05.046. PMID 18590827.
- Price, SA.; Bininda-Emonds, OR.; Gittleman, JL. (2005). "A complete phylogeny of the whales, dolphins and even-toed hoofed mammals – Cetartiodactyla". Biol Rev Camb Philos Soc. 80 (3): 445–473. doi:10.1017/s1464793105006743. PMID 16094808.
- Montgelard, C.; Catzeflis, FM.; Douzery, E. (1997). "Phylogenetic relationships of artiodactyls and cetaceans as deduced from the comparison of cytochrome b and 12S RNA mitochondrial sequences". Molecular Biology and Evolution 14 (5): 550–559. doi:10.1093/oxfordjournals.molbev.a025792. PMID 9159933.
- Spaulding, M.; O'Leary, MA.; Gatesy, J. (2009). "Relationships of Cetacea -Artiodactyla- Among Mammals: Increased Taxon Sampling Alters Interpretations of Key Fossils and Character Evolution". PLoS ONE 4 (9): e7062. Bibcode:2009PLoSO...4.7062S. doi:10.1371/journal.pone.0007062. PMC 2740860. PMID 19774069.
- Cetacean Species and Taxonomy. iucn-csg.org
- "The Society for Marine Mammalogy's Taxonomy Committee List of Species and subspecies".
- Perrin, W.F. (2012). "Eubalaena australis Desmoulins, 1822". World Cetacea Database. Retrieved 29 September 2012.
- Carwardine MH, Hoyt E (1998). Whales, Dolphins and Porpoises. Surry Hills, NSW: Reader's Digest. ISBN 0-86449-096-8.
- Müller, J. (1954). "Observations of the orbital region of the skull of the Mystacoceti" (PDF). Zoologische Mededelingen 32: 239–90.
- Branch, G.M., Branch, M.L, Griffiths, C.L. and Beckley, L.E. 2010. Two Oceans: a guide to the marine life of southern Africa ISBN 978-1-77007-772-0
- Department of Sustainability and Environment. 1998. Southern Right Whale - Eubalaena australis. Action Statement-Flora and Fauna Guarantee Act 1988. No.94. Retrieved on 31 October. 2014
- Whale and Dolphin Conservation Society. Endangered Species - SOUTHERN RIGHT WHALE Eubalaena australis. Retrieved on 31 October. 2014
- Crane, J. and R. Scott. (2002). "Eubalaena glacialis". Animal Diversity Web. Retrieved 27 June 2009.
- Waerebeek V.K., Santillán L., Suazo E. from The Peruvian Centre for Cetacean Research (2009). "ON THE NATIVE STATUS OF THE SOUTHERN RIGHT WHALE EUBALAENA AUSTRALIS IN PERU" (PDF). Boletín del Museo Nacional de Historia Natural, Chile 58: 75–82. Retrieved 2014-12-26.
- Fairfax Regional Media (30 July 2013). "White whale in Middleton". The Times. Retrieved 14 May 2015.
- Rice, Dale W. (1998). Marine mammals of the world: systematics and distribution. Society of Marine Mammalogy, Special Publication No. 4. ISBN 1891276034.
- Rosenbaum, H. C., R. L. Brownell Jr.; M. W. Brown C. Schaeff, V. Portway, B. N. White, S. Malik, L. A. Pastene, N. J. Patenaude, C. S. Baker, M. Goto, P. Best, P. J. Clapham, P. Hamilton, M. Moore, R. Payne, V. Rowntree, C. T. Tynan, J. L. Bannister and R. Desalle (2000). "World-wide genetic differentiation of Eubalaena: Questioning the number of right whale species" (PDF). Molecular Ecology 9 (11): 1793–802. doi:10.1046/j.1365-294x.2000.01066.x. PMID 11091315.
- Palaeobiology and Biodiversity Research Group, University of Bristol
- "List of Marine Mammal Species and Subspecies". Committee on Taxonomy. Society for Marine Mammology. 3 April 2012. Retrieved 29 September 2012.
- 1967fishdude. 2010. whale to close for comfort on YouTube. Re-retrieved on 27 November 2014
- BANKS A.,BEST P.,GULLAN A., GUISSAMULO5 A., COCKCROFT V., FINDLAY K. "Recent Sightings of Southern Right Whales in Mozambique" (PDF). Retrieved 3 April 2014.
- Baker S. (2012). Paternity study of right whales finds local fathers most successful. Oregon State University. Re-retrieved on 27 November 2014
- Turismo Miramar ARG.. 2011. 28 August. BALLENAS EN MIRAMAR. YouTube. Retrieved on 16 December 2014
- Kenney, Robert D. (2008). "Right Whales (Eubalaena glacialis, E. japonica, and E. australis)". In Perrin, W. F.; Wursig, B.; Thewissen, J. G. M. Encyclopedia of Marine Mammals. Academic Press. pp. 962–69. ISBN 978-0-12-373553-9. Retrieved 20 May 2012.
Kiribatiwas invoked but never defined (see the help page).
Cite error: The named reference
- Right Whale News, May 1998. Retrieved 24 July 2007.
- Macpherson D., 2014. First whale of the season spotted in Ballito. http://www.ecr.co.za/post/first-whale-of-the-season-spotted-in-ballito/. East Coast Radio. retrieved on 20-05-2014
- Umdloti Beach Accommodation. Umdloti Beach-Whale Sightings Register. http://www.umdloti.org/whales.html. retrieved on 20-05-2014
- The Namibian Sun. 2013. Southern right whale - The right whale to protect. Retrieved on 29 October 2014
- Travel News Namibia. 2012. The return of the whales. Retrieved on 29 October 2014
- Roux J.P.. Braby R.. Best B.P.. 2013. Movements of southern right whales between Namibia and South Africa - recolonization or populationexpansion - Does disappearance mean extirpation? - The case of Namibian rightwhales. SC/65a/ForInfo22. Retrieved on 26 October 2014
- Duke University (2008). "Spatial Ecology of the North Atlantic Right Whale (Eubalaena Glacialis)." (book). The ProQuest. Retrieved 2015-10-16.
- Richards R.. 2009. Past and present distributions of southern right whales (Eubalaena australis). New Zealand Journal of Zoology. Vol. 36: 447-459. 1175-8821 (Online); 0301-4223 (Print)/09/3604–0447. The Royal Society of New Zealand. Retrieved on 18 December 2014
- "HUMPBACK WHALE MIGRATION - Southern Right Whale (Eubalaena australis)". The Dolphin Encountours Research Center. Retrieved 2015-10-22.
- Report of the IWC Workshop on the Assessment of Southern Right Whales. SC/64/Rep5
- Camilah Antunes Zappes (1 March 2013). "The conflict between the southern right whale and coastal fisheries on the southern coast of Brazil". ResearchGate. Retrieved 14 May 2015.
- Report of the Scientific Committee; Panama City, Panama, 11–23 June, 2012 (PDF). Annual Meeting of the International Whaling Commission. Panama City, Panama: International Whaling Commission. 2012. pp. 48–49. IWC/64/Rep1Rev1. Retrieved 30 October 2012.
- "Southern right whale". WDC, Whale and Dolphin Conservation. Retrieved 14 May 2015.
- R. NATALIE P. GOODALL, BENEGAS G.L., DELLABIANCA N., RICCIALDELLI L., PIMPER E.L., 17 Jan 2014, The Presence of Southern Right Whales off Eastern Tierra del Fuego, 1987-2011.http://iwcoffice.co.uk/_documents/sci_com/workshops/SRW/S11-RW30.pdf. retrieved on 06-05-2014
- Ballena Franca navegó por la Ría. Retrieved on 16 December 2014
- Arias M.. 2009.Ballenas, en la ría de Puerto Deseado. The La Nación. Retrieved on 16 December 2014
- The Travel to Argentina. 2014. Whales off the coast of Mar del Plata. Retrieved on 16 December 2014
- The Visiting Argentina. 2014. Mar del Plata: Ballenas franca australes llegaron a sus costas. Retrieved on 16 December 2014
- Taringa!. Miramar: Avistaje frecuente de la ballena Franca Austral. Retrieved on 16 December 2014
- Isha S.. 2008. COMENTARIOS LLENOS DE AMOR! .... Retrieved on 3 January 2014
- "Uruguay in Photos - Whales in Punta". Uruguay in Photos. Retrieved 14 May 2015.
- Gubin A.. 2013. Ecoceanos: crítica sobrevivencia de las 50 ballenas Franca Austral chilenas. La Gran Época. Retrieved on 26 December 2014
- ObsChiloé CECPAN (2014). "ballena franca austral en Pumillahue, Chiloé.". p. YouTube. Retrieved 2015-03-09.
- "BALLENA AZUL -- Quienes Somos". Retrieved 14 May 2015.
- Gibbons J.. Capella J.J.. Kusch A.. Cárcamo j.. 2006. THE SOUTHERN RIGHT WHALE EUBALAENA AUSTRALIS (DESMOULINS, 1822) IN THE STRAIT OF MAGELLAN, CHILE. Anales Instituto Patagonia (Chile), 2006. 34: pp.75-80. Retrieved on 18 December 2014
- The Patagon _Journal. 2009. Southern Right Whale Spotted in Chilean Waters. Retrieved on 16 October 2014
- The Fundación Cethus. 2014. Realizamos nuestra 5° campaña en Cabo Vírgenes, Santa Cruz. Retrieved on 16 December 2014
- The Fundación Cethus. 2011. Cuarta campaña para el estudio de la Ballena franca austral en Cabo Vírgenes. Retrieved on 16 December 2014
- Gastón C (2013). "Eubalaena australis". Flickr. Retrieved 2015-03-09.
- Vernazzani G.B., Cabrera E., ROBERT L., Jr.Brownell L.R. (2013). "Eastern South Pacific southern right whale photo-identification catalog reveals behavior and habitat use patterns" (PDF). MARINE MAMMAL SCIENCE (the Society for Marine Mammalogy and Cetacean Conservation Center and National Marine Fisheries Service). doi:10.1111/mms.12030. Retrieved 2015-03-09.
- "Head of the Bight". Retrieved 14 May 2015.
- Southern right whales - South Australia 2009. YouTube. 15 August 2009. Retrieved 14 May 2015.
- South Australian Whale Centre, Latest Sightings Accessed 12 March 2015.
- Southern Right Whale added to Endangered Species List. YouTube. 6 May 2012. Retrieved 14 May 2015.
- Chased by a friendly whale -- a stand up paddle (SUP) experience on the NSW south coast, Australia. YouTube. 6 September 2010. Retrieved 14 May 2015.
- Whales at Moruya River II. YouTube. 26 August 2010. Retrieved 14 May 2015.
- Moruya. YouTube. 28 August 2010. Retrieved 14 May 2015.
- Narooma 130913 WTF. YouTube. 3 October 2013. Retrieved 14 May 2015.
- "Conservation Management Plan for the Southern Right Whale - A Recovery Plan under the Environment Protection and Biodiversity Conservation Act 1999 (2011–2021)" (PDF). Retrieved 2015-02-13.
- "This week at the station". Retrieved 14 May 2015.
- Nichols, Daphne (2006). Lord Howe Island Rising. Frenchs Forest, NSW: Tower Books. ISBN 0-646-45419-6. Retrieved on 3 November 2014
- Reid A.. Halderen V.L.. 2013. Wildlife Management - Impact of Deep Sea Oil Development on New Zealand marine wildlife. Department of Zoology. University of Otago. Retrieved on 18 December 2014
- Patenaude, Nathalie J. "Sightings of southern right whales around ‘mainland’ New Zealand" (PDF). Department of Conservation, Wellington, NZ. Retrieved May 2013.
- "Fiordland Coastal Newsletter" (PDF). Department of Conservation, Invercargill, NZ. Retrieved May 2013.
- "Southern right whales - something really special: Media release 22 October 2009". Doc.govt.nz. 22 October 2009. Retrieved 16 June 2012.
- "Otago 'hot spot' for whale sightings - Otago Daily Times". Rebecca Fox. 7 April 2013. Retrieved 9 April 2013.
- "Southern whales the right stuff for new era - Whales and Whaling - NZ Herald News". 188.8.131.52. Retrieved 16 June 2012.
- Stewart R., Todd B., 2001 A note on observations of southern right whales at Campbell Island, New Zealand. Journals of Cetacean Research Management Special Issue 2, pp.117–120, 2001. Retrieved on 9 July 2014
- "Kermadecs - Whales & Dolphins". Retrieved 14 May 2015.
- "Chatham Islands marine mammals". Retrieved 14 May 2015.
- "Southern right whale expedition 2009: Media release 1 September 2009". Doc.govt.nz. 1 September 2009. Retrieved 16 June 2012.
- Carroll L.E.. 2011. RETURN OF THE RIGHT WHALE: ASSESSMENT OF ABUNDANCE, POPULATION STRUCTURE AND GENEFLOW IN THE NEW ZEALAND SOUTHERN RIGHT WHALE. University of Auckland. Retrieved on 31 October 2014
- Miller E.C. (2007). "Current State of Knowledge of Cetacean Threats, Diversity, and Habitats in the Pacific Islands Region" (pdf). Whale and Dolphin Conservation Society Australasia. p. 44. Retrieved 2015-09-07.
- The RIGHT WHALE NEWS. 1999. Volume 6. Number 4. Retrieved on 8 December 2014
- Tristan da Cunha Government and the Tristan da Cunha Association. "Cetacea - Whales and Dolphins around the Tristan da Cunha Islands". Retrieved 5 April 2014.
- M. Postma, M. Wege, M.N. Bester, D.S. van der Merwe (2011). "Inshore Occurrence of Southern Right Whales (Eubalaena australis) at Subantarctic Marion Island". Retrieved 2015-03-10.
- Gaskin, D.E. (July 1964). "Return of the Southern Right Whale (Eubalaena australis Desm.) to New Zealand Waters, 1963". Tuatara 12 (2): 115–118. Retrieved 22 July 2007.
- Tonnessen, J. N. and A. O. Johnsen (1982). The History of Modern Whaling. United Kingdom: C. Hurst & Co. ISBN 0-905838-23-8.
- Tormosov D.D., Mikhaliev Y.A., Best P.B., Zemsky V.A., Sekiguchi K., Brownell R.L. (November 1998). "Soviet catches of southern right whales Eubalaena australis, 1951-1971. Biological data and conservation implications (abstract)". Biological Conservation 86 (2): 185–197. doi:10.1016/S0006-3207(98)00008-1. Retrieved 22 July 2007.
- Reeves, Randall R., Brent S. Stewart, Phillip J. Clapham and James. A Powell (2002). National Audubon Society: Guide to Marine Mammals of the World. United States: Alfred A. Knopf, Inc. ISBN 0-375-41141-0.
- Berzin A., Ivashchenko V.Y., Clapham J.P., Brownell L.R. Jr. (2008). "The Truth About Soviet Whaling: A Memoir" (PDF). DigitalCommons@University of Nebraska - Lincoln. Retrieved 2013-10-15.
- Department of Conservation. The conservation of whales in the 21st century - Whale diversity in New Zealand waters. Retrieved on 5 November 2014
- Petrobras, Projeto Baleia Franca. http://www.baleiafranca.org.br/ More information on Brazilian right whales is available in Portuguese
- "Appendix I" of the Convention on the Conservation of Migratory Species of Wild Animals (CMS). As amended by the Conference of the Parties in 1985, 1988, 1991, 1994, 1997, 1999, 2002, 2005 and 2008. Effective: 5 March 2009.
- "CMS Pacific Cetaceans MOU for Cetaceans and their Habitats in the Pacific Islands Region". Retrieved 14 May 2015.
- Increased harassment of Right Whales (Eubalaena australis) by Kelp Gulls (Larus dominicanus) at Península Valdés, Argentina. Rowntree, V.J., P. MacGuiness, K. Marshall, R. Payne, J. Seger, and M. Sironi, 1998. Marine Mammal Science. 14(1): 99 - 115. doi:10.1111/j.1748-7692.1998.tb00693.x
- Gulls' vicious attacks on whales. BBC News, 24 June 2009.
- "Ocean Alliance". Retrieved May 2013.
- "Uruguay se convirtió en un santuario de ballenas y delfines". Ecología. Cromo.com.uy. Retrieved 5 September 2013.
- "Els Vermeulen". Retrieved 14 May 2015.
- La Opinión Austral. 2012. Ballena franca en Deseado. Retrieved on 16 December 2014
- Lopes M.. 2013. TRES BALLENAS FRANCAS VISITARON LA RESERVA PROVINCIAL RIA DESEADO. Retrieved on 16 December 2014
- lopes M.. 2012. Puerto Deseado recibió a una ballena franca austral. Retrieved on 16 December 2014
- Acercando Naciones. Unbelievable: whales appeared off the coast of Mar del Plata. Retrieved on 16 December 2014
- PILAR ROSASCO FOTOGRAFIA. 2012. Las ballenas francas llegaron a Mar del Plata. The Hostnews Contenidos. Retrieved on 16 December 2014
- Diario Cuatro Vientos . 2014. Investigarán en Miramar la presencia de ballenas. Retrieved on 16 December 2014
- Roberto Álvarez R.. 2012. Enjoy your visit!. Department Of Tourism and Culture MGA.. Retrieved on 16 December 2014
- F5NOTICIAS.COM . 2012. Volvieron las ballenas a la ciudad. Retrieved on 16 December 2014
- Ballena franca retorna a Estrecho de Magallanes y abre nueva opción de avistamientos turísticos. Retrieved on 16 December 2014
- "Southern right whale back in NZ waters". TVNZ. Retrieved 14 May 2015.
- "First known NZ whale birth since end of hunting". Stuff. Retrieved 14 May 2015.
- "Rare whale sighting off Auckland coast". Retrieved 14 May 2015.
- Antarctica's Southern Right Whales. YouTube. 26 March 2012. Retrieved 14 May 2015.
- "Auckland Islands Marine Mammal Sanctuary". Retrieved 14 May 2015.
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- An online educational documentary film about southern right whales - whale trackers