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Squalius carolitertii 03 by-dpc.jpg
Squalius carolitertii
Squalius pyrenaicus 03 by-dpc.jpg
Squalius pyrenaicus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Cypriniformes
Family: Cyprinidae
Subfamily: Leuciscinae
Genus: Squalius
Bonaparte, 1837
Type species
Leuciscus squalus
Bonaparte, 1837

Cephalopsis Fitzinger, 1873
Iberocypris Doadrio, 1980
Microlepis Bonaparte, 1846

Squalius is a genus of fish in the family Cyprinidae found in Europe and Asia.[1] Hybridization is not rare in the Cyprinidae, including this genus. S. alburnoides is known to be of ancient hybrid origin, with the paternal lineage deriving from a prehistoric species related to Anaecypris; the latter mated with ancestral S. pyrenaicus. Present-day S. alburnoides mates with sympatric congeners of other species.[2]


There are currently 51 recognized species in this genus:


  1. ^ Froese, Rainer, and Daniel Pauly, eds. (2016). Species of Squalius in FishBase. October 2016 version.
  2. ^ Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.
  3. ^ a b Turan, D., Kottelat, M. & Doğan, E. (2013): Two new species of Squalius, S. adanaensis and S. seyhanensis (Teleostei: Cyprinidae), from the Seyhan River in Turkey. Zootaxa, 3637 (3): 308–324.
  4. ^ a b c d e f g h i j Özuluğ, M. & Freyhof, J. (2011): Revision of the genus Squalius in Western and Central Anatolia, with description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22 (2): 107-148.
  5. ^ Esmaeili, H.R., Khaefi, R. & Zamanian Nejad, R. (2016): Historical review on the taxonomy of Squalius berak Heckel, 1843 (Teleostei: Cyprinidae). FishTaxa, 1 (3): 118-126.
  6. ^ Khaefi, R., Esmaeili, H.R., Sayyadzadeh, G., Geiger, M.F. & Freyhof, J. (2016): Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4169 (1): 145–159.