|Reconstructed skull at the Museum für Naturkunde, Berlin|
Galton & Sues, 1983
Galton & Sues, 1983
Stygimoloch (meaning "Styx devil" in Latin) is a genus of pachycephalosaurid dinosaur from the end of the Cretaceous period, roughly 66 million years ago. It is controversial as only juvenile specimens have been discovered, and thus the possibility has been raised that it represents a juvenile Pachycephalosaurus. This hypothesis has been supported by numerous studies since, the most recent of which agreeing with its synonymy with Pachycephalosaurus. It is currently known from the Hell Creek Formation, Ferris Formation, and Lance Formation of the Western Interior (United States), where it lived alongside other Hell Creek taxa such as Tyrannosaurus, Triceratops, and Edmontosaurus.
It is a relatively large pachycephalosaur, with the skull being about 46 centimeters long (18 in). Among North American pachycephalosaurs, only Pachycephalosaurus is larger. Unlike other pachycephalosaurs, the domed skull is relatively small, slightly flattened from side to side, and pear-shaped; even when isolated this unusual dome can easily be distinguished from the broader, larger domes of Pachycephalosaurus. While the dome is reduced in size, the ornamentation over the skull is more elaborate than in any other pachycephalosaur. Short, conical hornlets covered the nose, and the back corners of the skull bore an enormous pair of massive, backward-pointing spikes, up to 5 centimeters in diameter (2 in) and 15 centimeters long (6 in); these are surrounded by two or three smaller spikes. The function of this unusual ornamentation is unknown. Even if other pachycephalosaurs did butt heads (which is a subject of continuing debate), the small dome of Stygimoloch suggests that this behavior was not as important. Instead, the skull ornament might have functioned for display, may have been used for self-defense, or perhaps were locked together and used in shoving matches, like the horns of deer. Another possibility is that the squamosal horns were used to inflict pain during flank-butting.
History of discovery
The type species, S. spinifer, was described by British vertebrate paleontologist Peter Galton and German paleontologist Hans-Dieter Sues of the National Museum of Natural History in 1983. Stygimoloch is characterized by clusters of spikes on the back of the skull, in which a long central horn is surrounded by 2-3 smaller hornlets, and a tall, narrow dome. Dracorex has similar but shorter hornlets and lacks a dome, and has been interpreted as either a distinct species, or a juvenile or female of Pachycephalosaurus or Stygimoloch. It has also been proposed that Stygimoloch represents partially grown individuals of Pachycephalosaurus, an interpretation that has been supported multiple times since.
The generic epithet Stygimoloch comes from the river Styx, which is said to run through the underworld in Greek Mythology, in reference to the Hell Creek Formation, and from Moloch, the name of a Canaanite god associated with child sacrifice, chosen in reference to the bizarre appearance of the animal. The word spinifer translates to "thorny", therefore the specific epithet translates to "thorny devil from the river Styx".
The pachycephalosaur Dracorex may actually be an individual of Stygimoloch or Pachycephalosaurus in which the dome and horns are not well-developed, either because the animal was a juvenile or because the animal was a female. This consideration was supported at the 2007 annual meeting of the Society of Vertebrate Paleontology. Jack Horner of Montana State University presented evidence, from analysis of the skull of the single existing Dracorex specimen, that this dinosaur may well be a juvenile form of Stygimoloch. In addition, he presented data that indicates that both Stygimoloch and Dracorex may be juvenile forms of Pachycephalosaurus. Horner and M.B. Goodwin published their findings in 2009, showing that the spike/node and skull dome bones of all three 'species' exhibit extreme plasticity, and that both Dracorex and Stygimoloch are known only from juvenile specimens while Pachycephalosaurus is known only from adult specimens. These observations, in addition to the fact that all three forms lived in the same time and place, lead them to conclude that Dracorex and Stygimoloch were simply juvenile Pachycephalosaurus, which lost spikes and grew domes as they aged. A 2010 study by Nick Longrich and colleagues also supported the hypothesis that all flat-skulled pachycephalosaurs were juveniles, suggesting that flat-skulled forms like Goyocephale and Homalocephale represent juveniles of dome-skulled adults.
Evans et. al. (2015) recognized the synonymy of Stygimoloch with Pachycephalosaurus when assigning new cranial material from the Scollard Formation of Alberta to cf. Pachycephalosaurus. Goodwin and Evans (2016) presented an analysis of the youngest ontogimorphs of Pachycephalosaurus known, and found that the unique features of Stygimoloch and Dracorex represent consistent morphological characters on an ontogenetic continuum of Pachycephalosaurus. As such, Stygimoloch and Dracorex represent junior synonyms of Pachycephalosaurus. No study has since contested this analysis or found these results incorrect, thus Stygimoloch most likely represents juvenile Pachycephalosaurus remains.
- Switek, Brian (December 7, 2007). "Lumping Dinosaurs: Stygimoloch a juvenile Pachycephalosaurus?". scienceblogs.com.
- "Quién es el Stygimoloch?". Dinosaurios.org. Retrieved 9 December 2013.
- Carpenter, Kenneth (1997). "Agonistic behavior in pachycephalosaurs (Ornithischia:Dinosauria): a new look at head-butting behavior". Contributions to Geology. 32 (1): 19–25.
- Galton, P. M. and H. D. Sues (1983). "New data on pachycephalosaurid dinosaurs (Reptilia: Ornithischia) from North America." Canadian Journal of Earth Sciences 20: 462-472.
- Goodwin, M. B., E. A. Buchholtz, et al. (1998). "Cranial anatomy and diagnosis of Stygimoloch spinifer." Journal of Vertebrate Paleontology 18(2): 363-375.
- Sues, H. D., & Galton, P. M. (1987). Anatomy and classification of the North American Pachycephalosauria (Dinosauria: Ornithischian). Palaeontographica Abt. A., 198: 1-40.
- Erik Stokstad,"SOCIETY OF VERTEBRATE PALEONTOLOGY MEETING: Did Horny Young Dinosaurs Cause Illusion of Separate Species?", Science Vol. 18, 23 Nov. 2007, p. 1236;
- Horner, John; Goodwin, Mark (October 27, 2009). "Extreme Cranial Ontogeny in the Upper Cretaceous Dinosaur Pachycephalosaurus". PLOS.
- Horner J.R. and Goodwin, M.B. (2009). "Extreme cranial ontogeny in the Upper Cretaceous Dinosaur Pachycephalosaurus." PLoS ONE, 4(10): e7626. Online full text
- Longrich, N.R., Sankey, J. and Tanke, D. (2010). "Texacephale langstoni, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA." Cretaceous Research, . doi:10.1016/j.cretres.2009.12.002
- Evans, David C.; Vavrek, Matthew J.; Larsson, Hans C. E. (2015-04-22). "Pachycephalosaurid (Dinosauria: Ornithischia) cranial remains from the latest Cretaceous (Maastrichtian) Scollard Formation of Alberta, Canada". Palaeobiodiversity and Palaeoenvironments. 95 (4): 579–585. doi:10.1007/s12549-015-0188-x. ISSN 1867-1594.
- Goodwin, Mark B.; Evans, David C. (2016-02-06). "The early expression of squamosal horns and parietal ornamentation confirmed by new end-stage juvenilePachycephalosaurusfossils from the Upper Cretaceous Hell Creek Formation, Montana". Journal of Vertebrate Paleontology. 36 (2): e1078343. doi:10.1080/02724634.2016.1078343. ISSN 0272-4634.
- TEDx talk by Jack Horner on shape-shifting dinosaur skulls and dinosaur misclassification.