Tiger versus lion
Historically, the comparative merits of the tiger (Panthera tigris) versus the lion (Panthera leo) have been a popular topic of discussion by hunters, naturalists, artists and poets, and continue to inspire the popular imagination in the present day. In the past, lions and tigers reportedly competed in the wilderness, where their ranges overlapped in Eurasia. The most common reported circumstance of their meeting is in captivity, either deliberately or accidentally.
- 1 Temperament
- 2 History in captivity
- 3 Coexistence in the Eurasian wilderness
- 4 Observed fights
- 5 Physical comparison
- 6 Professional opinions
- 7 Arts and literature
- 8 See also
- 9 Notes
- 10 References
- 11 Further reading
- 12 External links
Both the lion and tiger have fearsome reputations in their areas, whether amongst prey, sympatric predators, or people. In particular, both may prey on humans, though rates of man-eating tend to be higher for the tiger than for the lion. General differences in behaviour are listed below:
- The lion is a highly social animal and the tiger is a solitary animal. It is generally agreed that the tiger is the faster, smarter and more ferocious of the two so keepers of captive tigers must take care to avoid a sudden attack.
- Lions may roam in prides of up to 30 individuals headed by a mature male or group of related males. Male lions are typically either killed or pushed away by incoming male leadership. The majority of single roaming lions tend to be males preparing for maturation and assimilation with a new or existing pride. While male lions are generally larger and stronger than female lions, it is the close-knit female pride alliance that typically hunts and provides for the pride. By contrast, tigers are often solitary, socializing at other times. During mating, a male tiger and tigress would come together, and when together, they would behave more ferociously to other creatures, or even humans, if nearby, but the same would apply to lions.
More specifically, however, the Asiatic lion has similarities and differences with both its African relative and the tiger. For example, Asiatic lions are social like their African relatives, and females may practise promiscuity. However, the structures of the prides of African and Asiatic lions vary, with male Asiatic lions usually associating with females during times of mating, similar to tigers, and whereas Asiatic lionesses and tigresses may practise promiscuity to defend their cubs, African lionesses are believed not to do it for that purpose.
History in captivity
In the circuses of Ancient Rome, exotic beasts, including Barbary lions and tigers, were commonly pitted against each other. A mosaic in the House of the Faun in Pompeii shows a fight between a lion and a tiger. There are different accounts of which of these animals beat or killed the other, throughout time. Although lions and tigers can be kept together in harmony in captivity, conflicts between the two species in captivity, which ended up in fatalities, have also been recorded.
Coexistence in the Eurasian wilderness
Currently, India is the only country on Earth confirmed to have both lions and tigers in its wilderness. For now, they do not share the same territory in India, but they did in the past, and there is a project mentioned below that could make their meeting in the wild possible, if implemented.
Before the end of the 20th Century, lions and tigers had also occurred in other Asian or Eurasian nations, including Iran. As such, there is a word for 'Lion', which can also mean 'Tiger', and is used in Iran, South Asia and other areas, that is 'Sher' or 'Shir' (Persian: شیر), and its significance is discussed below. The lion is also of cultural importance to the Far East, including the land of the Indochinese tiger, but according to authors such as Reginald Innes Pocock (1939) and Nowell and Jackson (1996), the lion did not naturally occur there.
According to Colin Tudge (2011), given that both cats hunt large herbivores, it is likely that they had been in competition in Asia. Despite their social nature, lions might have competed with tigers on an individual basis, as they would with each other.
Apart from the possibility of competition, there are legends of Asiatic lions and tigers interacting together to produce hybrid offspring, which would be ligers or tigons. From the fossil record, besides genetics, it would appear that the modern lion and tiger were present in Eurasia since the Pleistocene, when now-extinct relatives also existed there. Additionally, in the days before Indian Independence, the Maharaja of Gwalior introduced African lions into his area, which is a habitat for Bengal tigers.
Asiatic lion and Bengal tiger
In India, or in the extended modern sense, the Subcontinent, Asiatic lions and Bengal tigers coexisted in a number of places, before the end of the 20th century. Amongst the places where they coexisted, or were reported to have been at least, are the area of Gwalior in the center, that of Mount Abu in the north, and that of Bahawalpur in what is now Pakistan. Kailash Sankhala (1978) said that the habitat and prey of the Indian lion was not like those of an African savannah, but like habitats of the Indian tiger to an extent, including the dry, deciduous Aravali part of Sariska Tiger Reserve in the State of Rajasthan, and that it was a difficult place for predators to hunt as groups.
Today, lions are found in Gir Forest National Park and surrounding areas in the region of Saurashtra, State of Gujarat, and tigers are found in other places, like Sariska Tiger Reserve and Ranthambore National Park in neighboring Rajasthan, the neighboring states of Madhya Pradesh and Maharashtra, and the Bengali Sunderbans. Gir, Sariska and Ranthambore are in the same ecoregion, that of Kathiawar-Gir dry deciduous forests. Though the Bengal tiger is reportedly extinct in northern and southeastern Gujarat, the nearest population of tigers to Kathiawar Peninsula is in the border-area of Gujarat, Maharashtra and Madhya Pradesh. It is located across the Gulf of Khambhat from Kathiawar Peninsula, and includes the Dangs' Forest and Shoolpaneshwar Wildlife Sanctuary. Either big cat can be called 'Sher' (Hindi: शेर) and have a fearsome reputation in the Subcontinent, and emigrate from its protected habitat.
The possibility of conflict, between lions and tigers, had been raised in relation to India's Asiatic Lion Reintroduction Project, which was meant to introduce Gir Forest's lions to another reserve which is considered to be within the former range of the Indian lion, that is Kuno Wildlife Sanctuary in Madhya Pradesh, before December 2017. Kuno was reported to contain some tigers that came from Ranthambore Park, including one called 'T-38'. Concerns were raised that the co-presence of lions and tigers would "trigger frequent clashes." The University of Minnesota's Lion Research Project describes one reason to delay the introduction of lions to Kuno Palpur, is the fear that tigers living there would kill the incoming lions. In a one-on-one encounter, it is believed that a Bengal tiger could beat an Indian lion, given its weight advantage. Despite the fact that the habitats of Indian lions and tigers are similar means that they both live in conditions that favor solitary hunters of prey, these lions are social like their African relatives, and may form fighting groups, whereas tigers are usually solitary, and it is believed that a group of lions (2 – 3 males) or lionesses (2 – 4 females) is more than match for a single tiger or tigress (see § Temperament). Therefore it would appear that in order for Asiatic lions to survive in an area with Bengal tigers, the lions would have to be translocated there as intact groups rather than as individuals.
Reginald Innes Pocock (1939) mentioned that some people had the opinion that the tiger played a role in the near-extinction of the Indian lion, but he dismissed this view as 'fanciful'. According to him, there was evidence that tigers inhabited the Subcontinent, before lions. The tigers likely entered Northern India from the eastern end of the Himalayas, through Burma, and started spreading throughout the area, before the lions likely entered Northern India from Balochistan or Persia, and spread to places like the Bengal and the Nerbudda River. Because of that, before the presence of man could limit the spread of lions, tigers reached parts of India that lions did not reach. However, the presence of tigers throughout India did not stop the spread of lions there, in the first place, so Pocock said that it is unlikely that Bengal tigers played a role, significant or subordinate, in the near-extinction of the Indian lion, rather, that man was responsible for it, as was the case with the decline in tigers' numbers. As such, Pocock thought that it was unlikely that serious competition between them regularly occurred, and that even if Indian lions and tigers met, the chance that they would fight for survival was as good as the chance that they would choose to avoid each other, and that their chances of success, if they were to clash, were as good as each other's.
Asiatic lion and Caspian tiger
Before the start of the 21st century, Afghanistan, Iran, Iraq, Syria, Turkey, and former members of the Union of Soviet Socialist Republics, such as Azerbaijan, were reported to have had Asiatic lions and Caspian tigers. Besides Asia, lions and tigers had occurred in Europe, in the region of the Black Sea, with tigers occurring in Ciscaucasia, and lions occurring in the Balkans, up to Thrace and Macedonia, and possibly the Danube River, at least.
Velikiy Kniaz Vladimir II Monomakh of Kievan Rus', in his work, "Poucheniya Detyam" (1117), said that while he ruled Turov (in what is now Belarus) and Chernigov (in what is now the Ukraine), he was on a hunt when he was attacked by a lyuti zver (Russian: лютый зверь, Old Russian for "fierce animal"). The zver sprang towards his thighs, and hurt him and his horse. Traditionally, the zver was considered to be a wolf or lynx, but, according to Heptner and Sludskii (1972), neither would spring at a rider or injure a horse, so it was more likely to be a big cat, with some people thinking that it could have been a leopard, or that it was more likely to be a tiger than a lion. The occurrence of the lion at the southern Russian Steppes, or the area of the mouth of the Don River, is disputed, whereas tigers likely occurred in the Russian Steppes or at the estuary of the Don River.
In Afghanistan, it is possible that lions occurred at least in the southwest and southern parts. Tigers bred at the upper reaches of the Hari Rud or Tedzhen Darya at Herat. Tigers were found at a tributary of the Amu Darya called the "Pyandzh River," from where they could invade another place (like Persian tigers that invaded what was the Soviet Union), and the Geri, Kunduz and Murghab Rivers. Intrusions from the Soviet Union were reported in the 1960s. The last known sighting of a tiger was in the Babatag Range, which is on the border between Afghanistan and Tajikistan, in 1998.
In ancient times, the Persian lion lived throughout much of its namesake home, including in northern regions near the Transcaucasian and Turkestani parts of the Soviet Union, which is why Heptner and Sludskii could not deny that they had been in the Turkestani region also. Many years ago, in the north, lions had been in the area of Tehran, and the Persian upland. Around the year 900, they were encountered in the south, although not frequently. However, in the 1870s, they occurred in western region, in the southwestern part of the Zagros Mountains, near Mesopotamia, and in forested areas which were south and southeast of Shiraz. Persian tigers also occurred in regions close to the Soviet Union, including the north-western region, enough for them to invade the Trans-Caucasus and Turkestan, including those of the Atrek Basin and Gorgan.
The Euphrates and Tigris Rivers flow from Turkey to Iraq, through the Syrian region. Lions were seen along the upper reaches of the Euphrates (Biledzhik, 1877; Alston and Danford, 1880) in the 1870s, before disappearing there by the end of the 19th Century, though they otherwise survived in that period, in Mesopotamia and the Arabian peninsula. In the 1850s, lions also occurred in the upper courses of the Tigris, near Mosul in the north. In the 1860s, there were many lions in reed marshes, along the banks of the two rives, though mostly in their lower reaches (Blanford, 1876). Two lions in the region of Mosul were reported for the last time in 1914, and the last lion in Iraq was slaughtered on the lower Tigris in 1918. Mazandaran tigers also occurred in Iraq, on the banks of the Euphrates and Tigris. In 1887, a tiger was killed near Mosul (Kock, 1990).
In the Syrian region, before the end of the 19th Century, the lion occurred from what is now the country of Syria in the north, to the area of Palestine in the south. In particular, lions had been present in the area of Aleppo, even as late as 1891 (Kinnear, 1920). Up to the 1970s, the tiger had been reported in the area of Hatay, which includes Amuq Valley, and had been transferred from Syria to Turkey, during the Second World War. Turkish tigers were also reported from the Plain of Selçuk in the western region, to sparse forests and riverine corridors in the eastern region, which borders Iraq and Syria. In February 1970, a tiger was reportedly killed near Uludere in Şırnak Province, Hakkari Region. Anatolian lions had been in parts of the eastern region, apparently up to the 19th Century. Before that, the lion had been throughout Asia Minor, excepting the region of the Pontic Mountains in the north.
In what was to be the Transcaucasian Socialist Federative Soviet Republic in the Soviet Union in 1922, in the northern part of its range there, bypassing the eastern part of the Caucasus, the lion occurred in an area which extended unevenly from foot-hills and the Araks River near Yerevan (in what is now Armenia) in the east, almost to Tbilisi (in what is now Georgia) in the west, from Absheron Peninsula (in what is now Azerbaijan) in the south, to the Samur River (in the region of what is now the border between Azerbaijan and Dagestan in Russia) in the north. This place includes what are now the Lagodekhi Protected Areas in the Caucasus mixed forests' ecoregion. Lions were hunted by local hunters called 'shirvans' or 'shirvanshakhs', and became extinct by the end of the 10th Century. The Hyrcanian tiger was found in the areas of Tbilisi and Baku in Apsheron Peninsula (which has a wildlife sanctuary in the Caspian Hyrcanian mixed forests' ecoregion), and was reported to have intruded territories, like those of Baku and Tbilisi, from other places, like that of the Talysh Mountains and Lankaran Lowland in what is now Azerbaijan. The Trans-Caucasus is home to a tugay type of forest, and lions and tigers would have hunted prey like deer here.
For what used to be the Turkestani region of the Soviet Union, which now comprises the countries Kazakhstan, Kyrgyzstan, Turkmenistan and Uzbekistan, Heptner and Sludskii could not exclude the possibility of lions occurring in the southern part of the area, which is close to Iran, and along the upper Amu Darya, which has a tributary called "Sherabad Darya," which touches a town called 'Shirabad'. Doctor A. B. Meyer, in his book, The Antiquity of the Lion in Greece, said that the lion did indeed occur in the region of Khaurism, between Afghanistan and the Caspian Sea. This was based on Abbott's 1834 book, A Narrative of a journey from Heraut to Khiva, Moscow and St. Petersburgh, which said that the predators of Khaurism included both the lion and tiger. In the southwestern part of Turkestan, tigers occurred in the area of the Kopet Dag, along the Atrek River to the Caspian Sea, and the river's tributaries, the Chandyr and Sumbar Rivers, including the area of Tedzhen, often as intruders from Iran. They also occurred in the regions of the Amu and Syr Daryas, and others, in a vast area extending to the region of Western Siberia or Lake Baikal in the east, where the Amur tiger also reportedly occurred. Two tigers that were captured in southwestern Tajikistan harbored tapeworms (Taenia bubesei) which were also recovered from the lion, according to Chernyshev (1953).
Cave lion and tiger
In prehistoric times, before the 10th millennium BC, the Upper Pleistocene Eurasian cave lion had occurred throughout much of Eurasia, including the Caucasus, what is now Siberia in Russia, and what used to be Beringia. Judging from cave paintings, such as those of Chauvet, cave lions were social, having formed hunting groups like their modern relatives. Their great distribution may have enabled them to influence the distributions of sympatric carnivores like the tiger, through direct or indirect competition. In fact, during the early Pleistocene, tigers, including the prehistoric Wanhsien tiger, appear to have been confined to the Far East, from Siberia in the north to the Sunda Islands in the south. It was during the late Pleistocene or Holocene, around when the cave lion became rare or extinct, that the tiger spread westwards to places like the Caucasus and Indian Subcontinent.
In addition to historical recordings, clashes between lions and tigers were reported or even caught on camera, in the 19th and 20th centuries. It was not always clear which species regularly beat the other, according to Doctor Packer (2015).
- Roman Emperor Titus had Bengal tigers compelled to fight African lions, and the tigers always beat the lions.
- In July 1808, Sylvanus Urban said that Mr Bolton had a friend who claimed to have seen a fight between the lion and tiger at a circus in Verona. Though the tiger had attacked first, it yielded to the stronger lion.
- In 1830, a tiger attacked a lion at a menagerie in Turin, Rome. Despite having attacked first, the lion got it on its back, and used its jaws to hold the tiger's throat. The tiger died after that.
- Clark (1838) said that a British officer, who resided many years at Sierra Leone, saw many fights between lions and tigers, and that the tiger 'universally' won.[a]
- In 1857, an 18-month-old tiger at the Bromwich Zoo broke into the cage of an adult lion. The pair fought, and the young tiger ripped the lion's stomach. The lion died minutes later.
- According to Porter (1894), a tiger called 'Gunga', which belonged to the King of Oude, killed thirty lions, and destroyed another after being transferred to the zoological garden in London.
- According to the Gettysburg Compiler and The Baltimore Sun (1899), towards the end of the 19th century in India, the Gaekwad of Baroda, that is Sayajirao III, arranged a fight in an amphitheater, between a Barbary lion called 'Atlas', from the Atlas Mountains between Algeria and Morocco, and a Bengal tiger from the Indian region of Shimla, both large and hungry (with their diets reduced before the fight), before an audience of thousands, instead of between an Indian lion and the tiger, as Indian lions were believed to be no match for Bengal tigers.[b] The tiger was more than 10 feet (3.0 metres) long, over 4 feet (120 centimetres) feet at the shoulder, had strong shoulders and long teeth and claws, and was agile. The lion looked taller at the head than the tiger, and had a large mane, legs and paws. The tiger was seen as "the personification of graceful strength and supple energy," whereas the lion was seen as the "embodiment of massive power and adamantine muscle." In the fight, both cats sustained injuries, and although the tiger sometimes retreated from Atlas, it would come back to fight it, and in the end, managed to scratch Atlas to death, though Atlas pushed it off in one final move, before dying. The Gaekwad agreed to pay 37,000 rupees, accepted that the tiger was the "King of the Cat Family," decreed that Atlas' body be given a Royal burial, and that the tiger should have a "cage of honour" in the menagerie of Baroda, and decided to prepare the tiger for a battle with a Sierran Grizzly bear weighing more than 1,500 lb (680 kilograms). The battle was to happen after the tiger recovered from its wounds.
- At the Coney Island animal show in 1909, a performing lion attacked a chained tiger by leaping through the air, landing on the tiger's back. Though hampered by the heavy neck chain fastened to the iron bars of the arena, the tiger was more than a match for the lion and mangled it to death.
- In May 1914, at New York's Bronx Zoo, the barrier between the cages of Rajah, an 8-year-old Bengal tiger, and Huerte, a Nubian lion which was 2 or 3 years old, and had been sick for some time, got opened in an "unaccountable manner." For the first few minutes, the more agile Huerte appeared to be winning, but when it aimed for its injured opponent's neck, about an hour into the fight, Rajah aimed for its nape. Not only did Rajah manage to bite Huerte's nape, but it also broke Huerte's back, thus slaughtering it.
- In 1934, a fully grown African lion killed a mature Bengal tiger a short time after these circus animals were unloaded from the train, before trainers could separate them.
- Bert Nelson (1938) said that in Chicago, when 20 lions and tigers were mixed together for an act at a circus, a fight occurred, lasting for about 10 minutes. No fatalities were mentioned, but Nelson said that order was 'restored' when tigers used escape doors to flee.
- At South Perth Zoo, 1949, in a three-minute fight between a lion and a tiger, the lion killed the tiger. The fight occurred when the tiger put his head through a connecting slide. The lion caught the tiger by the throat, and, dragging it through the opening, killed it before the keepers arrived.
- By 21 February 1951, Beatty had seen 50 tigers killed by lions throughout his career.
- In September 1951, at a corporation zoo in Madras, India, an eighteen-year-old tiger called 'Vikram' entered the cage of a seven-year-old lion called 'Leo', and got into a fight with it. Badly mauled, including in the femur, Vikram retreated into its cage, and despite receiving medical attention, died.
- An Indian Prince organised a filmed fight in a deep pit in the compound of his palace. The lion had killed the tiger, according to Kailash Sankhala (1978).
- In March 2011, a Bengal tiger at the Ankara Zoo passed through a gap, between its cage and that of a lion, and killed it with a single paw swipe. "The tiger severed the lion's jugular vein in a single stroke with its paw, leaving the animal dying in a pool of blood," officials said.
In the wilderness
- It appears from the 15th-century book Anvâr-i Suhaylî (Persian: اَنوارِ سُهيلى, "Lights of the Canopus") that the lion and tiger competed for dominance in the region of the Tigris River, that is Mesopotamia.
- Herne (1855) mentioned that in the Indian jungle between the village of Elaw, city of Baroche, and Gulf of Cambay, north of the city of Surat and its Ghauts, about 6.0 or 7.0 mi (9.7 or 11.3 km) from the village, he and his party, which included locals, heard a tiger's roar. Pursuing it, they caught a glimpse of it, but by that time, the tiger had attacked a local. It disappeared with the victim, and after pursuing it for about 50.0 yd (150.0 ft), they heard the roar of lion, and besides it, sounds which suggested that it was in a struggle with the tiger, such as growls. The party not only managed to see the lion and tiger rolling about in their battle, after going through bushes, but also the man who fell victim to the tiger. The author termed both the lion and tiger as "tyrants of the forest," given that they would attack weaker creatures. The tiger was about the same size as the lion, but more agile. As for the lion, it used greater strength, and its mane, which was somewhat deeper than those of its bigger African cousins, could protect its head from the tiger's claws, though not other parts of its body, such as the back. They were as determined and brave as each other, but the lion endured. It caught the tiger's throat, turned it on its back, and killed it by clawing its abdomen open. The lion was thus hailed as the "King of Beasts." Otherwise, the fight had been harsh for both beasts, to the extent that the author felt that it would avenge their victims.
- The Sun (New York) reported that in a depopulated Indian village at the bank of a creek connected to the Cauvery River, about 30.0 mi (48.3 km) north-west of Bangalore, a hunter injured by a venomous creature saw a tiger on his left-hand side, and a lion on his right-hand side. The tiger was a "rousing big fellow, who had seen 15 years of his life," and had muscular limbs. The male lion was "medium-sized." Both of them stalked him, but they did not notice each other at first as they were separated by a wall that was about 4.0 ft (120 cm) tall, and their focus was on the witness. When they got closer to him, the tiger scented the lion, and behaved like an angry cat, which included making a noise that startled the latter. The lion showed its teeth in response, and after reaching the end of the wall, roared at its foe. After the lion's head showed around the wall, the crouching tiger pounced on it, and rolled over with it. As they fought, which involved making bites or scratches, they growled in a way that turned the hunter's hair gray, in his own words. Tigers often kill victims by biting their throats, and keeping their hold on them for as long as necessary, but that was not the case with this struggle. Despite different descriptions of their sizes by the narrator, and that the tiger was more agile than the lion, the tiger's neck was vulnerable to a bite by the lion, and for reasons like these, it was difficult for either cat to defeat the other, overall. After they temporarily retreated from each other, the hunter could see that they were both injured. Still, they were determined to destroy each other. The lion and tiger respectively roared and snarled. The narrator suspected that their hatred for each other may have been because both had been hunting him at the same time, therefore, their respective presences interfered with each other's hunt for him. The tiger pounced on the lion's back, rolling over or falling with it again, and struggling to its feet like it. The lion seemed helpless as the tiger held onto its fore shoulder, before making a move in which it managed to catch the tiger's neck. Now, the tiger seemed helpless, before making a move to use its hind claws to force the lion to release its hold on it. Though the tiger was the aggressor this time, their struggle became more like that of dogs unable to beat each other. They bled from nose to tail as they moved away from the witness, towards the creek. They fell into the water, which was about 2.0 ft (61 cm) deep, and this stopped the fight. They retreated from each other, limping into the forest.[c]
- Rivalry between the Asiatic lion and "Siberian tiger"[d] is mentioned in Hamilton M. Wright's work, in The San Francisco Call (1911).
Generally, in terms of dimensions of the body and weight, the modern tiger and lion are the two largest species of the genus Panthera and the cat family. Variations in opinions and measurements exist for them, especially across different populations or subspecies. Apart from that, similarities and differences exist for other characteristics, such as the lengths of their skulls. As for their prehistoric relatives, such as the Ngandong tiger and American lion, they were considered to have been rather large, but measurements, estimates or opinions for their weights or sizes differ.
Favoring the tiger
- John Varty, owner of the Londolozi Reserve in South Africa, said, "People always ask me which one is bigger? If a tiger and a lion had a fight, which one would win? Well, I've seen tigers crunch up a full-grown leopard tortoise like it was nothing. And lions try, but they just don't get it right. If there's a fight, the tiger will win, every time."
- The animal rescue organisation Big Cat Rescue of Tampa, Florida answered, "While it would depend on the size, age, and aggressiveness of the specific animals involved, generally tigers have a significant advantage."
- The conservation charity Save China's Tigers stated "Recent research indicates that the tiger is indeed stronger than the lion in terms of physical strength. Lions hunt in prides, so it would be in a group and the tiger as a solitary creature so it would be on its own. A tiger is generally physically larger than a lion. Most experts would favor a Siberian and Bengal tiger over an African lion."
- John Smith Clarke, a British lion tamer, said, in a lecture on the fight between a tiger and a lion given to the Glasgow Zoological Society, while showing the actual fight on the screen, "in 100 cases out of 100 the tiger would always beat the lion. It was far more agile, it was not so clumsy in its movements, it was equally strong, it was equally armed, but it fought in a different way. The tiger very often fought rolling on its back and held the lion in its grip until it defeated him."
- The BBC (2016), in a 3-round study of agility, strength and intelligence, favored the tiger in the case of intelligence, due to the time spent for a particular challenge.
- National Geographic Channel's documentary The Last Lions of Asia mentioned that a Bengal tiger had a weight advantage of 50 kilograms (110 lb) over an Asiatic lion, and could kill a lone lion in a fight.
Favoring the lion
- Clyde Beatty, the animal trainer and performer who owned several tigers, lions, hyenas, and other exotic animals, believed that in nine out of ten times, "a full-grown lion would whip a full-grown tiger". He mentioned that since he first began mixing the animals, 25 of his tigers (two of them being Siberian tigers, the rest of them being Bengal tigers) were killed in the circus arena, but there was not a single lion casualty.
- Renowned naturalist and conservationist of India, Kailash Sankhala wrote in his book Tiger that the tiger would be unable to get close to lion's vital joints because of his thick mane, and that the tiger would be vulnerable to the lion. He mentioned that once an Indian prince organized a fight in which the lion killed the tiger, and opined that "a tiger is no match for even single lion of equal strength".
- Carl Hagenbeck, a trainer from Hamburg, said that the lion and tiger were alike in "good temper and reliability." He cited the example of a lion being trained for a month to do tricks, and of a tiger taking five weeks to do so.
Arts and literature
Battles between the two were painted in the 18th and 19th centuries by Eugène Delacroix, George Stubbs and James Ward. Ward's paintings, which portrayed lion victories in accordance with the lion's symbolic value in Great Britain, have been described as less realistic than Stubbs.
The British Seringapatam medal shows a lion defeating a tiger in battle; an Arabic language banner on the medal displays the words Asad Allāh al-Ghālib (Arabic: أسَـد الله الـغَـالِـب, "Lion of God the Conqueror"). The medal commemorated the British victory at the 1799 Battle of Seringapatam (in the town now known as Srirangapatna) over Tipu Sultan — who used tigers as emblems, as opposed to the British emblematic use of lions.
Lion and Tiger Fighting by James Ward, 1797
The Garden Arbor (German: Die Gartenlaube) by Ernst Keil's Nachfolger, 1858
Painting by Friedrich Specht, 1883
Sculptures of a tiger and lion fighting, with the former dominating the latter, by Emile-Joseph-Alexandre Gouget, in Le musée d'Art et d'Industrie de la ville de Roubaix
The Seringapatam medal shows a lion defeating a tiger
18th-century naturalists and authors compared the species' characters, generally in favor of the lion. Oliver Goldsmith ranked the lion first among carnivorous mammals, followed by the tiger, which in his view "seems to partake of all the noxious qualities of the lion, without sharing any of his good ones. To pride, courage and strength, the lion joins greatness, clemency and generosity; but the tiger is fierce without provocation, and cruel without necessity." Charles Knight, writing in The English Cyclopaedia, disparaged the opinions of naturalists Georges-Louis Leclerc, Comte de Buffon and Thomas Pennant in this context, stating "the general herd of authors who eulogise the 'courage, greatness, clemency and generosity' of the lion, contrasting it with the unprovoked ferocity, unnecessary cruelty and poltroonery of the tiger, becomes ridiculous, though led by such names as Buffon and Pennant."
- English literature compared their battle strengths. The poets Edmund Spenser, Allan Ramsey, and Robert Southey described lion victories. In the view of a 19th-century literary critic, these contests established "sovereignty of the animal world."
- According to John Hampden Porter (1894, page 176), Lockington said that the jaguar reigned 'supreme' as the "terror of the forest," and that this applied to the lion and tiger in the 'desert' and 'jungle', respectively.
In the Hindu epic Mahabharata, Narada told Srinjaya that tigers were fiercer and more ruthless than lions. This is in contrast with other literature from ancient India, which prefers the lion to the tiger. For example, Vedic literature depicted the lion, rather than the tiger, as the "king of the forest."
The lion and tiger rival each other in Iranian literature. For example, Humphreys and Kahrom, in their 1999 book Lion and Gazelle: The Mammals and Birds of Iran, treated them as the "two greatest and most beautiful" of Iranian carnivores, albeit being extinct there. As with the lion, the tiger's Persian name was used for people and places.
The lion's mane
About the lion's mane, Knight wrote "The lion has owed a good deal to his mane and his noble and dignified aspect; but appearances are not always to be trusted." In fact, a study was done by scientists Craig Packer and Peyton West that claimed that the mane of the lion was strictly for mating purposes. Darker-maned lions were more often picked by females to breed, while light-maned lions were not so lucky. A lion's mane did not always purposely help in a fight, and it might even hinder the male lion, slowing it down when it attacks, according to Packer and West.
However, Dereck Joubert (1996), who spent many hours studying lions in Botswana, believed that the mane could defend the lion's throat, in a fight with another lion, and Clyde (1939) and Kailash (1978) believed that the mane could defend part of the lion's neck, in a fight against the tiger. Reports also support these claims, for example, when the Shimlan tiger fought Atlas, it tried biting Atlas' neck, but it could not, due to the mane blocking its teeth, and interfering with its respiratory system. According to Heptner and Sludskii (1997), Barbary and Cape lions had the "most luxuriant and extensive manes" among lions, with "tresses on flanks and abdomen." As for the Indian lion, which has a smaller mane than its African cousins, apart from African lions that have weak manes or are maneless, during the fight reported by The Sun (New York) (1889), though the tiger could bite the lion's body, it was not mentioned to have bitten the lion's neck.
In Paalai, a Tamil film, there is dialogue about the characteristics of the tiger and lion. It concludes that the tiger is superior. In the film, the tiger is the symbol and flag of the native Tamil tribal people and the lion is the symbol and flag of non-Tamil Singhal (literally meaning 'Leonine') people.
- Animal Face-Off
- Atlas the Barbary lion versus the Bengal tiger of Simla
- Lions, Tigers and Bears
- The Jeff Corwin Experience, Episode 10 — India: Between the Tiger and the Lion
- Tyavarekoppa Lion and Tiger Safari
- As a West African nation, Sierra Leone would have had indigenous lions, but not tigers.
- For more details, see Asiatic lion versus Bengal tiger.
- As reported by St. Landry Democrat (1887), The Eaton Democrat (1887), The Iola Register (1887), The Milan Exchange (1887), and The Sydney Mail (1889), in chronological order.
- Though this term was used, genetic tests showed that the Amur tiger was closely related to the Caspian tiger, which also inhabited the former Soviet Union (where Siberia is located), and whose range would have overlapped with that of the lion in places like the Caucasus.
- "Bannerghatta National Park". Bengaloorutourism.com. Retrieved 2014-05-24.
- Tudge, C. (2011). Engineer In The Garden. Random House. p. 42. ISBN 9781446466988.
- Heptner, V. G.; Sludskii, A. A. (1992) . Mlekopitajuščie Sovetskogo Soiuza. Moskva: Vysšaia Škola [Mammals of the Soviet Union, Volume II, Part 2]. Washington DC: Smithsonian Institution and the National Science Foundation. pp. 83–202. ISBN 90-04-08876-8.
- Nowell, Kristin; Jackson, Peter (1996). Wild Cats: Status Survey and Conservation Action Plan (PDF). Gland, Switzerland: IUCN/SSC Cat Specialist Group. pp. 17–149. ISBN 2-8317-0045-0.
- José Ortega y Gasset (2007). Meditations on Hunting. ISBN 978-1-932098-53-2.
- John Hampden Porter (1894). Wild beasts; a study of the characters and habits of the elephant, lion, leopard, panther, jaguar, tiger, puma, wolf, and grizzly bear. pp. 76–256. Retrieved 2014-01-19.
- Ronald Tilson, Philip J. Nyhus (2010), "Tiger morphology", Tigers of the world, Academic Press, ISBN 9780815515708
- William Bridges (22 August 1959). Lion vs. tiger: who'd win?. The Spokesman-Review. Retrieved 2016-02-28.
- Lion against tiger. The Baltimore Sun. 26 January 1899. p. 3.
- Thomas, Isabel (2006). Lion vs. Tiger. Raintree. ISBN 978-1-4109-2398-1.
- London Express (27 December 1900). "Animal Fighters". Past Papers. Retrieved 2017-01-09.
- Packer, C. "Frequently asked questions". University of Minnesota Lion Research Project. Retrieved 28 June 2011.
- "A Terrible Struggle". The Sydney Mail. 21 December 1889. Retrieved 2016-12-29.
- Humphreys, P., Kahrom, E. (1999). Lion and Gazelle: The Mammals and Birds of Iran. Images Publishing, Avon.
- Clyde Beatty, Earl Wilson (1941), Jungle performers
- Roland Auguet (1994). Cruelty and civilization: the Roman games. ISBN 978-0-415-10453-1.
- "Lion and Tiger Fight to Death, Lion is Victor". Logansport Press. 16 November 1934. Retrieved 16 November 2014.
- Brakefield, Tom (1993). Big Cats: Kingdom of Might. Voyageur Press. p. 44. ISBN 978-0-89658-329-0.
- Kirk Bates (28 February 1951). "When a Lion fights a Tiger". The Milwaukee Journal.
- Pocock, R. I. (1939). The Fauna of British India, including Ceylon and Burma. Mammalia. – Volume 1. Taylor and Francis Ltd., London. Pp. 197–222.
- Schaller, p. 142.
- Sunquist, M.; Sunquist, F. (2002). Wild Cats of the World (1st ed.). Chicago: University Of Chicago Press. pp. 7–350. ISBN 978-0-22-677999-7.
- Joslin, P. (1973). The Asiatic lion: a study of ecology and behaviour. University of Edinburgh, UK: Department of Forestry and Natural Resources.
- Meena V. (2008). Reproductive strategy and behaviour of male Asiatic Lions. Dehra Dun: Wildlife Institute of India.
- Sharma, N. (6 October 2016). "Promiscuity of mother lioness saes cubs in Gir: Study". Hindustan Times. Retrieved 2017-09-21.
- "Barbary Lion Information". Beinglion.com. Retrieved 2016-06-29.
- "Barbary Lion - Panthera leo leo". The Sixth Extinction Website. Archived from the original on 3 March 2016. Retrieved 2016-06-29.
- Hoage, Robert J., Roskell, Anne and Mansour, Jane, "Menageries and Zoos to 1900", in New World, New Animals: From Menagerie to Zoological Park in the Nineteenth Century, Hoage, Robert J. and Deiss, William A. (ed.), Johns Hopkins University Press, Baltimore, 1996, pp.8-18. ISBN 0-8018-5110-6
- Anthony King (2002). The natural history of Pompeii. Cambridge University Press. ISBN 978-0-521-80054-9.
- "Tiger, lion and bear form unusual friendship". 7 December 2009.
- Beatty, Clyde (Oct 1939). "Which is the King of Beasts": 563. Retrieved 16 November 2014.
- Herbert Treadwell Wade (1930). The New International Encyclopaedia, Volume 22. Dodd, Mead and Company. Retrieved 2017-05-29.
- "Lion against tiger". Gettysburg Compiler. 7 February 1899. Retrieved 2016-02-28.
- Charles Frederick Partington (1835). "Felis, the cat tribe". The British cyclopæedia of natural history. Orr & Smith.
- Shankaranarayanan, P.; Banerjee, M.; Kacker, R. K.; Aggarwal, R. K. & Singh, L. (1997). "Genetic variation in Asiatic lions and Indian tigers" (PDF). Electrophoresis. 18 (9): 1693–1700. doi:10.1002/elps.1150180938. PMID 9378147. Archived from the original (PDF) on 23 July 2013.
- Tales of travellers. Tales of travellers; or, A view of the world. p. 453. Archived from the original on 1838.
- Karanth, K. U. (2003). "Tiger ecology and conservation in the Indian subcontinent". Journal of the Bombay Natural History Society. 100 (2–3): 169–189. Archived from the original on 10 March 2012.
- "Lion vs Tiger". YouTube. 8 July 2008. Retrieved 4 February 2012.
- Preparations for the reintroduction of Asiatic lion Panthera leo persica into Kuno Wildlife Sanctuary, Madhya Pradesh, India by A.J.T. Johnsingh, S.P. Goyal, Qamar Qureshi; Cambridge Journals Online; Oryx (2007), 41: 93-96 Cambridge University Press; Copyright 2007 Fauna & Flora International; doi:10.1017/S0030605307001512; Published online by Cambridge University Press 05Mar2007
- "Tigers moving from Rajasthan to Madhya Pradesh, officials concerned". Times of India. 19 April 2013. Retrieved 2016-02-07.
- Haas, S.K.; Hayssen, V.; Krausman, P.R. (2005). "Panthera leo" (PDF). Mammalian Species. 762: 1–11. doi:10.1644/1545-1410(2005)762[0001:PL]2.0.CO;2. Archived from the original (PDF) on 28 July 2017.
- "Asiatic lion". Cat Specialist Group. Archived from the original on 25 August 2010. Retrieved 2016-06-28.
- Mazák, V. (1981). "Panthera tigris" (PDF). Mammalian Species. 152: 1–8. doi:10.2307/3504004. Archived from the original (PDF) on 9 March 2012.
- Luo, S.-J.; Kim,J.-H.; Johnson, W. E.; van der Walt, J.; Martenson, J.; Yuhki, N.; Miquelle, D. G.; Uphyrkina, O.; Goodrich, J. M.; Quigley, H. B.; Tilson, R.; Brady, G.; Martelli, P.; Subramaniam, V.; McDougal, C.; Hean, S.; Huang, S.-Q.; Pan, W.; Karanth, U. K.; Sunquist, M.; Smith, J. L. D.; O'Brien, S. J. (2004). "Phylogeography and genetic ancestry of tigers (Panthera tigris)". PLoS Biology. 2 (12): e442. doi:10.1371/journal.pbio.0020442. PMC . PMID 15583716. Retrieved 2016-03-21.
- "Kipling's list of names in the stories", excerpted from volume XII of The Complete Works, Sussex edition, 1936.
- Shahbazi, Shapur A. (2001). "Flags (of Persia)". Encyclopaedia Iranica. 10. Retrieved 2016-03-10.
- Goswamy, B. N. (2002). "Where does the Lion come from in ancient Chinese culture? Celebrating with the Lion Dance". The Tribune Newspaper, Chandigarh, India. Retrieved 2010-12-14.
- Upham, E. (1829). The History and Doctrine of Budhism: Popularly Illustrated: with Notices of the Kappooism, Or Demon Worship, and of the Bali, Or Planetary Incantations, of Ceylon. London: R. Ackermann.
- "Image 5 of 20". myanmar-image.com. Retrieved 2010-09-17.
- Ray, Nick; Daniel Robinson; Greg Bloom (2010). Cambodia. Lonely Planet. p. 99. ISBN 978-1-74179-457-1.
- "Liger". messybeast.com. Retrieved 2016-06-21.
- ligerfacts.org. "The Liger - Meet the World's Largest Cat". Retrieved 2016-07-17.
- Description of ligers at Bestiarium.kryptozoologie.net
- Description of ligers at Lairweb.org.nz
- Shi, Wei (2005). Growth and Behaviour: Epigenetic and Genetic Factors Involved in Hybrid Dysgenesis. Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology. 11. Uppsala: Acta Universitatis Upsaliensis. p. 9. ISBN 91-554-6147-6.
- Kitchener, A. C.; Dugmore, A. J. (2000). "Biogeographical change in the tiger, Panthera tigris". Animal Conservation. 3 (2): 113–124. doi:10.1111/j.1469-1795.2000.tb00236.x.
- Jhala, Y. V., Qureshi, Q., Sinha, P. R. (Eds.) (2011). Status of tigers, co-predators and prey in India, 2010. National Tiger Conservation Authority, Govt. of India, New Delhi, and Wildlife Institute of India, Dehradun. TR 2011/003 pp-302
- John McLeod, The history of India, page 1, Greenwood Publishing Group, 2002, ISBN 0-313-31459-4
Milton Walter Meyer, South Asia: A Short History of the Subcontinent, pages 1, Adams Littlefield, 1976, ISBN 0-8226-0034-X
Jim Norwine & Alfonso González, The Third World: states of mind and being, pages 209, Taylor & Francis, 1988, ISBN 0-04-910121-8
Boniface, Brian G.; Christopher P. Cooper (2005). Worldwide destinations: the geography of travel and tourism. Butterworth-Heinemann. ISBN 978-0-7506-5997-0.
Judith Schott & Alix Henley, Culture, Religion, and Childbearing in a Multiracial Society, pages 274, Elsevier Health Sciences, 1996, ISBN 0-7506-2050-1
Raj S. Bhopal, Ethnicity, race, and health in multicultural societies, pages 33, Oxford University Press, 2007, ISBN 0-19-856817-7
Lucian W. Pye & Mary W. Pye, Asian Power and Politics, pages 133, Harvard University Press, 1985, ISBN 0-674-04979-9
Mark Juergensmeyer, The Oxford handbook of global religions, pages 465, Oxford University Press US, 2006, ISBN 0-19-513798-1
Sugata Bose & Ayesha Jalal, Modern South Asia, pages 3, Routledge, 2004, ISBN 0-415-30787-2
- Kinnear, N. B. (1920). "The past and present distribution of the lion in south eastern Asia". Journal of the Bombay Natural History Society. 27: 34–39. Retrieved 2017-02-01.
- Jhala, Y. V.; Gopal, R.; Qureshi, Q., eds. (2008), Status of the Tigers, Co-predators, and Prey in India (PDF), TR 08/001, National Tiger Conservation Authority, Govt. of India, New Delhi; Wildlife Institute of India, Dehradun, archived from the original (PDF) on 2 June 2013
- "Kathiawar-Gir dry deciduous forests". Terrestrial Ecoregions. World Wildlife Fund. Retrieved 2017-01-29.
- Uncle Ray (8 May 1952). "Tigers And Bears Are Found In Himalayas". St. Petersburg Times. Retrieved 2017-05-29.
- D. K. Harshey & Kailash Chandra (2001). Mammals of Madhya Pradesh and Chhattisgarh. Zoos´ Print Journal 16(12): 659-668 online
- Negi, Sharad Singh (2002), Handbook of National Parks, Wildlife Sanctuaries and Biosphere Reserves in India (3rd Edition), Indus Publishing, p. 151, ISBN 978-81-7387-128-3
- Kailash, Sankhala (1978). Tiger: The Story Of The Indian Tiger. Collins. p. 119. ISBN 0002161249.
- "Asiatic Lion population up from 411 to 523 in five years". Desh Gujarat. 10 May 2015. Retrieved 2016-11-26.
- "Gir National Park & Wildlife Sanctuary". Govt. of Gujarat. Forests and environment Dept. Archived from the original on 22 November 2015. Retrieved 2016-01-11.
- "Gujarat Map". mapsofindia.com. Retrieved 2016-03-21.
- "India wild tiger census shows population rise". BBC News. 28 March 2011. Retrieved 2011-02-08.
- Significant bird records and local extinctions in Purna and Ratanmahal Wildlife Sanctuaries, Gujarat, India-PRANAV TRIVEDI and V. C. SONI
- "Mahal Eco Campsite". Gujarat Tourism. Retrieved 2017-01-25.
- "Vansda National Park". Gujarat Tourism. Retrieved 2017-01-29.
- Garshelis, David L.; Joshi, Anup R.; Smith, James L. D. & Rice, Clifford G. "Sloth Bear Conservation Action Plan" (PDF). Archived from the original (PDF) on 23 July 2011. Retrieved 18 April 2011.
- "Stalemate on translocation of Gir lions Kuno Palpur in Madhya Pradesh to be used as tiger habitat now". Hindustan Times. 7 December 2017. Retrieved 2018-01-27.
- Sharma, R. (5 December 2017). "Tired of Gujarat reluctance on Gir lions, MP to release tigers in Kuno". Times of India. Retrieved 2018-01-27.
- "MP not fit for Asiatic lions, Gujarat tells PM". Times of India. 19 June 2006. Retrieved 2011-06-28.
At a two-hour meeting of National Board of Wildlife presided by Prime Minister Manmohan Singh here, Govind Patel said the "presence of tigers in the Kuno Palpur sanctuary would trigger frequent clashes between the two carnivores over territories – tiger and lions—which can never co-exist in the same place."
- Sevruguin, A. (1880). "Men with live lion". National Museum of Ethnology in Leiden, The Netherlands; Stephen Arpee Collection. Retrieved 2018-03-26.
- Masseti, M. (2009). Carnivores of Syria In: E. Neubert, Z. Amr , S. Taiti, B. Gümüs (eds.) Animal Biodiversity in the Middle East. Proceedings of the First Middle Eastern Biodiversity Congress, Aqaba, Jordan, 20–23 October 2008. ZooKeys 31: 229–252.
- "The Extinction Website: Panthera tigris virgata". The Extinction Website. 11 April 2010. Archived from the original on 12 April 2016. Retrieved 2016-04-03.
- Jackson, P.; Nowell, K. (2008). "Panthera tigris ssp. virgata". IUCN Red List of Threatened Species. Version 2012.2. International Union for Conservation of Nature. Retrieved 2016-01-30.
- "Balkans". Encyclopædia Britannica. Retrieved 2016-04-10.
The Balkans are usually characterized as comprising Albania, Bosnia and Herzegovina, Bulgaria, Croatia, Kosovo, Macedonia, Montenegro, Romania, Serbia, and Slovenia—with all or part of each of those countries located within the peninsula. Portions of Greece and Turkey are also located within the geographic region generally defined as the Balkan Peninsula, and many descriptions of the Balkans include those countries too. Some define the region in cultural and historical terms and others geographically, though there are even different interpretations among historians and geographers....Generally, the Balkans are bordered on the northwest by Italy, on the north by Hungary, on the north and northeast by Moldova and Ukraine, and on the south by Greece and Turkey or the Aegean Sea (depending on how the region is defined) ...
- Sommer, R. S.; Benecke, N. (2006). "Late Pleistocene and Holocene development of the felid fauna (Felidae) of Europe: A review". Journal of Zoology. 269: 7. doi:10.1111/j.1469-7998.2005.00040.x.
- Canadian Institute of Ukrainian Studies (1988). "Kyivan Rus'". Encyclopedia of Ukraine. Retrieved 2016-03-29.
- Leonid Smilovitsky (2004). "A Byelorussian Border Shtetl in the 1920s and 1930s: The Case of Turov". Retrieved 2016-03-29.
- "Weatherbase: Historical Weather for Chernihiv, Ukraine". Weatherbase. Retrieved 2016-03-29.
- Jungius, H.; Chikin, Y.; Tsaruk, O.; Pereladova, O. (2009). Pre-Feasibility Study on the Possible Restoration of the Caspian Tiger in the Amu Darya Delta (PDF). WWF Russia. Retrieved 19 August 2016.
- Daoudy, Marwa (2005). Le Partage des Eaux entre la Syrie, l'Irak et la Turquie. CNRS. pp. 1–269. ISBN 2-271-06290-X. Retrieved 2016-04-06.
- Hatt, R. T. (1959). The mammals of Iraq. Ann Arbor: Museum of Zoology, University of Michigan.
- "The Caspian Tiger". lairweb.org.nz. Retrieved 2016-04-06.
- Johnson, K. (2002). "The Status of Mammalian Carnivores in Turkey". University of Michigan.
- Sekercioglu, Cagan; Sean Anderson; Erol Akcay; Rasit Bilgin; Ozgun Can; Gurkan Semiz (December 2011). "Turkey's Globally Important Biodiversity In Crisis" (PDF). Biological Conservation. 144 (12): 2752–2769. doi:10.1016/j.biocon.2011.06.025. Retrieved 3 October 2013.[permanent dead link]
- "The Soviet Period - History - Azerbaijan - Asia". Retrieved 2016-02-17.
- Hovasapyan, Zara (1 August 2012). "When in Armenia, Go Where the Armenians Go". Armenian National Committee of America. Retrieved 2016-02-06.
Made of local pink tufa stones, it gives Yerevan the nickname of "the Pink City."
- "Preliminary Results of 2014 General Population Census of Georgia" (PDF). NATIONAL STATISTICS OFFICE OF GEORGIA. Retrieved 2016-02-12.
- "Administrative, density and territorial units and land size by economic regions of Azerbaijan Republic for January 1. 2007". Archived from the original on 24 November 2007. Retrieved 12 February 2016.
- "Samur: Azerbaijan". National Geospatial-Intelligence Agency. 1995–2012. Retrieved 2016-02-17.
- Michael Kohn, John Noble and Danielle Systermans, Georgia, Armenia & Azerbaijan, 3rd ed. Footscray, Victoria / London: Lonely Planet, 2008, ISBN 9781741044775, p. 115.
- "Caucasus mixed forests". Terrestrial Ecoregions. World Wildlife Fund.
- "National Parks: Absheron National Park - Ministry of Ecology and Natural Resources of Azerbaijan". 26 October 2011. Archived from the original on 2011-07-26. Retrieved 2017-06-16.
- "Absheron National Park Official Website - Ministry of Ecology and Natural Resources of Azerbaijan". 1 September 2012. Archived from the original on 2012-09-01. Retrieved 2017-06-16.
- "Ecosystem Profile: Caucasus". Conservation International. Archived from the original on 25 July 2008. Retrieved 2017-07-26.
- Britannica, Encyclopædia (1978). "Turkistan". Encyclopædia. Retrieved 2016-02-01.
- Oxford Dictionaries, Oxford University Press Retrieved: 20 February 2016.
- Abbott, James (1856). A Narrative of a journey from Heraut to Khiva, Moscow and St. Petersburgh. 1. Khiva: James Madden. p. 26.
- Arduini, P. & Teruzzi, G. 1993. The MacDonald encyclopedia of fossils. Little, Brown and Company, London. 320pp.
- 'Supersize' lions roamed Britain BBC News 1 April 2009
- Hooijer, Dirk Albert.; 1947. Pleistocene remains of Panthera tigris (Linnaeus) subspecies from Wanhsien, Szechwan, China, compared with fossil and Recent tigers from other localities. American Museum Novitates; no. 1346. Accessed on 2013-08-22.
- "Valvert 2014_Wanhsien Tiger Data Sheet". Retrieved 2015-06-20.
- Yamaguchi, Nobuyuki; Cooper, Alan; Werdelin, Lars; MacDonald, David W. (August 2004). "Evolution of the mane and group-living in the lion (Panthera leo): a review". Journal of Zoology. 263 (4): 329–342. doi:10.1017/S0952836904005242.
- Turner, A. and Antón, M. (1997). The Big Cats and Their Fossil Relatives: An Illustrated Guide to Their Evolution and Natural History. Columbia University Press. ISBN 978-0-231-10228-5.
- The Medical times and gazette: A Journal of Medical Science. p. 626. Archived from the original on 1850.
- Urban, S. (1834). The Gentleman's Magazine, Volumes 156 – 157. F. Jeffries. Retrieved 2016-11-24.
- "Lion and tiger fight". Newcastle Morning Herald and Miners' Advocate. 31 October 1930. Retrieved 2016-11-24.
- The Descent of Man, and Selection in Relation to Sex. Charles Darwin
- "His Highness Sayajirao Gaekwad III". Gaekwadsofbardoa.com. Retrieved 2017-04-08.
- Lawson, Alastair (10 December 2011). "Indian maharajah's daring act of anti-colonial dissent". The BBC. Retrieved 2011-12-10.
- "Tiger whips Lion". Boston News Access. 7 March 2011. Retrieved 14 March 2011.
- "Lion Fights Tiger". The Horsham Times. 12 May 1914. Retrieved 2017-10-19.
- "Lion Fights Tiger: Latter Scores A Win". Warracknabeal Herald. 12 May 1914. Retrieved 2017-10-19.
- "Tiger and lion in death fight at New York Zoo". The Fairmont West Virginian. 8 May 1914. Retrieved 2017-07-10.
- "Lion and tiger in death battle". The Pittsburgh Press. 8 May 1914. Retrieved 2017-07-10.
- Rover, R. (15 January 1938). "Our Junior Section: Ralph Rover's Letter". The Age. Retrieved 2017-07-10.
- "Lion Kills Tiger". Examiner. 30 July 1949. Retrieved 16 November 2014.
- "Jungle Cats' Clash Not In Act". Newcastle Morning Herald and Miners' Advocate. 22 February 1951. Retrieved 2016-11-25.
- "Beatty fights off lion after tiger is killed". The Tuscaloosa News. 20 February 1951. Retrieved 2014-01-19.
- "ZOO TIGER SUCCUMBS: FOUGHT WITH LION". Retrieved 2016-06-29.
- "Tiger Kills Lion In Turkish Zoo". BBC News. 7 March 2011. Retrieved 14 March 2011.
- Eastwick, Edward B (transl.) (1854). The Anvari Suhaili; or the Lights of Canopus Being the Persian version of the Fables of Pilpay; or the Book Kalílah and Damnah rendered into Persian by Husain Vá'iz U'L-Káshifí. Hertford: Stephen Austin, Bookseller to the East-India College. Retrieved 2017-05-02.
- Herne, P. (1855). "XXIII: Domus. Surat. The nature of the jungles beyond. A boa constrictor. A tiger. A lion. Terrible conflict. A Banyan tree.". Perils and Pleasures of a Hunter's Life; or the Romance of Hunting by Peregrine Herne. Cornell University Library. pp. 194–204. Retrieved 2017-07-08.
- Schaller, G. (1967). The Deer and the Tiger: A Study of Wildlife in India. Chicago: Chicago Press.
- "An Awful Fight: A Combat Between a Tiger and a Lion". St. Landry Democrat. 30 April 1887. Retrieved 2017-04-08.
- "An Awful Fight: A Combat Between a Tiger and a Lion". The Eaton Democrat. 14 April 1887. Retrieved 2017-04-08.
- "An Awful Fight: A Combat Between a Tiger and a Lion". The Iola Register. 22 April 1887. Retrieved 2017-04-08.
- "An Awful Fight: A Combat Between a Tiger and a Lion". The Milan Exchange. 23 April 1887. Retrieved 2017-04-08.
- "When the Mastodon Walked Up Market Street". The San Francisco Call. 30 July 1911. Retrieved 2017-05-28.
- Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O’Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z.; Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News (Special Issue 11).
- Driscoll, C.A., Yamaguchi, N., Kahila Bar-Gal, G., Roca, A.L., Luo, S.-J., Macdonald, D. and O’Brien, S. J. (2009). "Mitochondrial phylogeography illuminates the origin of the extinct Caspian tiger and its relationship to the Amur tiger". PLOS ONE. 4: 1–8. Bibcode:2009PLoSO...4.4125D. doi:10.1371/journal.pone.0004125. PMC . PMID 19142238.
- Merriam, J. C. & Stock, C. 1932: The Felidae of Rancho La Brea. Carnegie Institution of Washington Publications 442, 1–231.
- DeSantis, L. R.; Schubert, B. W.; Scott, J. R.; Ungar, P. S. (2012). "Implications of diet for the extinction of saber-toothed cats and American lions". PLoS ONE. 7 (12): e52453. Bibcode:2012PLoSO...752453D. doi:10.1371/journal.pone.0052453. PMC . PMID 23300674.
- "John Varty Interview". Country Life. 10 October 2013. Archived from the original on 5 October 2013. Retrieved 10 October 2013.
- "Big Cat Rescue FAQ". Big Cat Rescue. 9 February 2012. Retrieved 9 February 2012.
- "Save China's Tigers Questions". Save China's Tigers. 9 November 2011. Archived from the original on 4 September 2011. Retrieved 9 November 2011.
- "Lion versus the tiger". The Glasgow Herald. 26 March 1937.
- "lion vs tiger:". BBC Earth Unplugged. 10 August 2016.
- Hornaday, W. T. (31 December 1905). "Mental capacity of Animals". The Evening Standard. Retrieved 2017-07-10.
- Frank McLynn (2006). 1759: The Year Britain Became Master of the World. Canongate Books. p. 163. ISBN 978-0-8021-4228-3.
George Stubbs, the most famous and original animal painter of his time who was just reaching his peak in 1759, liked to display combats of lion versus tiger, though he did not commit the egregious mistake made in James Ward's animal pictures painted later in the century where the lion symbolises Britain and the tiger India; in reality, as we know very clearly from the obscene animal fights staged by the Ancient Romans in the arena, the tiger would win every time
- Maya Jasanoff (2007). Edge of Empire: Lives, Culture, and Conquest in the East, 1750–1850. Random House. ISBN 978-0-307-42571-3.
- Charles Knight (1854). The English cyclopaedia: a new dictionary of Universal Knowledge. Bradbury and Evans. p. 219. Retrieved 30 June 2011.
- Oliver Goldsmith; Georges Léopold C.F.D. Cuvier (baron de.) (1847). A history of the earth and animated nature, with an intr. view of the animal kingdom tr. from the Fr. of Baron Cuvier, notes and a life of the author by W. Irving. p. 367. Retrieved 30 June 2011.
- Charles Francis Richardson (1883). Good literature: a literary eclectic weekly, Volume 5. AbeBooks. p. 114.
- Prof. Muneo Tokunaga John D. Smith K M Ganguli. "Tiger in Hindu epic Mahabharatha". The Mahabharata in Sanskrit. Sacred Texts. pp. SECTION LXVIII. Retrieved 2016-06-15.
- Mitra, S. (2005). Gir Forest and the saga of the Asiatic lion. New Delhi: Indus. ISBN 8173871833.
- "Mane of the Lion". Bigcatnews.blogspot.com. September 2006. Archived from the original on 25 April 2012. Retrieved 4 February 2012.
- Joubert, D. (18 May 1996). "Letters : By any other mane". New Scientist. Retrieved 2018-01-22.
- Schoe, Marjolein; Sogbohossou, Etotépé A.; Kaandorp, Jacques and de Iongh, Hans (2010) Progress Report – collaring operation Pendjari Lion Project, Benin. Funded by the Dutch Zoo Conservation Fund.
- Isabel Thomas (2006). Lion vs. Tiger. Raintree. ISBN 978-1-4109-2398-1.
- Jerry Pallotta; Rob Bolster (2009). Lion vs. tiger. Scholastic. ISBN 978-0-545-17571-5.