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Diplophysa dalaica.jpg
Triplophysa dalaica
Triplophysa stoliczkai Day cleaned.png
Triplophysa stolickai
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Cypriniformes
Family: Nemacheilidae
Genus: Triplophysa
Rendahl (de), 1933
Type species
Nemacheilus hutjertjuensis
Rendahl, 1933

Triplophysa is a genus of fish in the family Nemacheilidae found mainly in and around the Qinghai-Tibet Plateau in China.[1] Currently, the genus is a mixed assemblage of species. Some lineages have been identified and treated as subgenera (Hedinichthys, Indotriplophysa, Labiatophysa, Qinghaichthys and Tarimichthys), but following Kottelat and the Catalog of Fishes are treated as genera in Wikipedia.[2][3] FishBase, however, includes these in Triplophysa.[4]


Triplophysa zhaoi holds the record for the lowest altitude for Asian fish: it is found at 50 m (160 ft) below sea level in swamps of the Lükqün oasis, in the Turpan Depression in Xinjiang. In the other end, Triplophysa stolickai holds the record altitude for Asian fish: it is found at 5,200 m (17,100 ft) above sea level in hot springs near the Longmu Lake in western Tibet.[2] Triplophysa dalaica has been used as model species to study adaptation to high-altitude hypoxia and 13 positively selected genes involved in hypoxia response have been identified.[5] Some species are blind cave-dwellers.[1][6]


There are currently 131 recognized species in this genus:[2]


  1. ^ a b Chen, X.-Y. & Yang, J.-X. (2005): Triplophysa rosa sp. nov.: a new blind loach from China. Journal of Fish Biology, 66 (3): 599–608.
  2. ^ a b c Kottelat, M. (2012): Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). Raffles Bulletin of Zoology, Suppl. No. 26: 1-199.
  3. ^ Eschmeyer, W.N. (30 Aug 2016). "Catalog of Fishes". California Academy of Sciences. 
  4. ^ Froese, Rainer, and Daniel Pauly, eds. (2016). Species of Triplophysa in FishBase. June 2016 version.
  5. ^ Wang, Y., Yang, L., Wu, B., Song, Z. & He, S. (2015): Transcriptome analysis of the plateau fish (Triplophysa dalaica): Implications for adaptation to hypoxia in fishes. Gene, 565 (2): 211–220.
  6. ^ Romero, A., Zhao, Y. & Chen, X. (2009): The Hypogean fishes of China. Environmental Biology of Fishes, 86 (1): 211–278.
  7. ^ Wu, Y.-Y., Sun, Z.-Y. & Guo, Y.-S. (2016): A new species of the genus Triplophysa (Cypriniformes: Nemacheilidae), Triplophysa daochengensis, from Sichuan Province, China. Zoological Research, 37 (5): 290-296.
  8. ^ a b c Lan, J., Gan, X., Wu, T. & Yang, J. (2013). Cave Fishes of Guangxi, China. Science Press. p. 284. ISBN 9787030376589. 
  9. ^ Zheng, L.-P., Yang, J.-X. & Chen, X.-Y. (2012): A new species of Triplophysa (Nemacheilidae: Cypriniformes), from Guangxi, southern China. Journal of Fish Biology, 80 (4): 831–841.
  10. ^ Lin, Y., Li, C. & Song, J.-K. (2012): A new species of troglobitic loach (Cypriniformes, Balitoridae), Triplophysa jiarongensis, from Guizhou, China. Acta Zootaxonomica Sinica, 37 (3): 640–647.
  11. ^ a b Bashir, A., Bisht, B.S., Mir, J.I., Kumar, R. & Patiyal, R.S. (2016): Morphological, molecular characterization and taxonomic status of Triplophysa marmorata and Triplophysa kashmirensis (Cypriniformes: Nemacheilidae) from Kashmir valley, India. Revista de Biología Tropical, 64 (2): 473-482.
  12. ^ Wu, T.-J., Yang, J. & Lan, J.-H. (2012): A New Blind Loach Triplophysa lihuensis sp. nov. (Teleostei: Balitoridae) from Guangxi, China. Zoological Studies, 51 (6): 874–880.
  13. ^ Ren, Q., Yang, J.-X. & Chen, X.-Y. (2012): A new species of the genus Triplophysa (Cypriniformes: Nemacheilidae), Triplophysa longliensis sp. nov., from Guizhou, China. Zootaxa, 3586: 187–194.
  14. ^ Yang, J., Wu, T.-J. & Yang, J.-X. (2012): A new cave-dwelling loach, Triplophysa macrocephala (Teleostei: Cypriniformes: Balitoridae), from Guangxi, China. Environmental Biology of Fishes, 93 (2): 169–175.
  15. ^ He, C., Zhang, E. & Song, Z. (2012): Triplophysa pseudostenura, a new nemacheiline loach (Cypriniformes: Balitoridae) from the Yalong River of China. Zootaxa, 3586: 272–280.
  16. ^ Li, W.-J., Chen, X.-C. & Hu, Y.-P. (2015): A new species of the genus Triplophysa (Nemacheilinae), Triplophysa qilianensis sp. nov., from Qinghai, China. Zootaxa, 3905 (3): 418–424.
  17. ^ Yang, H.-F., Li, W.-X. & Chen, Z.-M. (2016): A new cave species of the genus Triplophysa from Yunnan, China. Zoological Research, 37 (5): 297-300.
  18. ^ Liu, S. W.; Pan, X. F.; Yang, J. X.; Chen, X. Y. (2017). "A new cave-dwelling loach, Triplophysa xichouensis sp. nov. (Teleostei Nemacheilidae) from Yunnan, China". Journal of Fish Biology. 90 (3): 834. doi:10.1111/jfb.13201. ISSN 0022-1112. PMID 28155227. 
  19. ^ Yan, S.-L.; Sun, Z.-Y.; Guo, Y.-S. (2015). "A new species of Triplophysa Rendahl (Cypriniformes, Nemacheilidae) from Sichuan Province, China". Zoological Research. 36 (5): 299–304. PMID 26452694.