Ukrainian stickleback

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Ukrainian stickleback
Ukrainian stickleback from the Dniester Estuary, Ukraine
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Gasterosteiformes
Family: Gasterosteidae
Genus: Pungitius
Species: P. platygaster
Binomial name
Pungitius platygaster
(Kessler, 1859)
  • Gasterosteus platygaster Kessler, 1859
  • Gasterosteus platygaster aralensis Kessler, 1877
  • Gasterosteus platygaster caucasicus Kessler, 1877
  • Gasterosteus platygaster danubica Steindachner, 1899
  • Gasterosteus platygaster kessleri Yakovlev, 1870
  • Gasterosteus platygaster niger Yakovlev, 1870
  • Gasterosteus pungitius niger Yakovlev, 1870
  • Pungitius platygaster aralensis (Kessler, 1877)
  • Pygosteus platygaster nuda Berg, 1905

The Ukrainian stickleback (Pungitius platygaster) also known as Caspian ninespine stickleback; southern ninespine stickleback; Aral ninespine stickleback, is a species of fish in the Gasterosteidae family. It is found in Afghanistan, Bulgaria, Iran, Kazakhstan, Moldova, Romania, Russia, Serbia, Montenegro, and Ukraine (Black Sea, Sea of Azov, Aral Sea and Caspian Sea basins).

A combination of two characters: lack of caudal peduncle keel, presence of large lateral scutes. It is similar to P. hellenicus with the following differences. Body darker than P. hellenicus (juvenile specimens from Iran show the same color as P. hellenicus). All the bones well ossified, cranial bones and pelvic bones highly sculptured. Pelvic girdle present with one spine and one small soft ray on each side. Dorsal spines 8-11, inclined alternatively to left and right. Last spine slightly longer than the others, which are relatively uniform. Dorsal soft rays 6-10; anal soft rays 6-9; gill rakers 9-11; bony scutes 29-32 with 7-12 large lateral scutes; total vertebrae 29-31; precaudal vertebrae usually 13. Caudal fin truncated.


Keivany, Y. 1996. Taxonomic revision of the genus Pungitius with emphasis on P. hellenicus. MSc thesis. Department of Biological Sciences, University of Alberta. Edmonton. 98 pp.

Keivany, Y., and J.S. Nelson. 2000. Taxonomic review of the genus Pungitius, ninespine sticklebacks (Teleostei, Gasterosteidae). Cybium, 24(2): 107-122.

Keivany, Y. and J.S. Nelson. 2004. Phylogenetic relationships of sticklebacks (Gasterosteidae), with emphasis on ninespine sticklebacks (Pungitius spp.). Behaviour, 141(11/12): 1485-1497.

World Conservation Monitoring Centre 1996.Pungitius platygaster. 2006 IUCN Red List of Threatened Species. Downloaded on 5 August 2007.