From Wikipedia, the free encyclopedia
Jump to navigation Jump to search

Upeneus parvus - Flickr - NOAA Photo Library.jpg
Upeneus parvus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Perciformes
Family: Mullidae
Genus: Upeneus
Type species
Mullus vittatus
Forsskål, 1775

Hypeneus Agassiz, 1846
Upeneoides Bleeker, 1849
Megalepis Bianconi, 1857
Pennon Whitley, 1941

Upeneus is a genus of goatfishes native to the Atlantic, Indian and Pacific oceans.


There are currently 37 recognized species in this genus:

Species[2] Taxon authority Common name Environment & range[3]
Upeneus asymmetricus Lachner, 1954 Asymmetrical goatfish Philippines, Indonesia
Upeneus australiae B. J. Kim & Nakaya, 2002 Australian goatfish Eastern Indian Ocean and Western Pacific; Australia
Upeneus davidaromi Golani, 2001 Darom's goatfish Red Sea; deep-water (150–500 m)
Upeneus doriae Günther, 1869 Gilded goatfish Persian Gulf and the Gulf of Oman
Upeneus filifer J. D. Ogilby, 1910 Pennant goatfish West-central Pacific
Upeneus francisi J. E. Randall & Guézé, 1992 Francis' goatfish Norfolk Island and New Zealand
Upeneus guttatus F. Day, 1868 Two-tone goatfish Indo-West Pacific: the Red Sea to South Africa and to northern Australia
Upeneus heemstra Uiblein & Gouws, 2014 [4] Heemstra goatfish Western Indian Ocean: Coast of Africa to India & Sri Lanka
Upeneus indicus Uiblein & Heemstra, 2010 Tall-fin goatfish Western India
Upeneus itoui M. Yamashita, Golani & Motomura, 2011 [5] Oriental goatfish Southern Japan
Upeneus japonicus Houttuyn, 1782 Japanese goatfish Japan north to Korea and south to Taiwan and Hong Kong
Upeneus lombok Uiblein & W. T. White, 2015 [6] Lombok goatfish Lombok, Indonesia
Upeneus luzonius D. S. Jordan & Seale, 1907 Dark-barred goatfish Pacific waters around the Philippines, Indonesia and northwestern Australia
Upeneus margarethae Uiblein & Heemstra, 2010 Margaretha's goatfish Indian Ocean
Upeneus mascareinsis Fourmanoir & Guézé, 1967 Mascarene goatfish Madagascar, Mozambique, Réunion, Western Indonesia
Upeneus moluccensis Bleeker, 1855 Goldband goatfish Indo-West Pacific, has invaded the Mediterranean through the Suez Canal[7]
Upeneus mouthami J. E. Randall & Kulbicki, 2006 Moutham's goatfish Western Central Pacific
Upeneus niebuhri Guézé, 1967 [4] Niebuhr’s goatfish Gulf of Suez
Upeneus nigromarginatus A. R. Bos, 2014 [8] Black-margined goatfish Philippines (Davao Gulf)
Upeneus oligospilus Lachner, 1954 Short-fin goatfish Persian Gulf
Upeneus parvus Poey, 1852 Dwarf goatfish Western Atlantic
Upeneus pori Ben-Tuvia & Golani, 1989 Por's goatfish Red Sea to southern Oman,has invaded the Mediterranean through the Suez Canal[9]
Upeneus quadrilineatus C. T. Cheng & C. X. Wang, 1963 Four-stripe goatfish Southern Japan to southern China
Upeneus randalli Uiblein & Heemstra, 2011 Persian Gulf
Upeneus saiab Uiblein & Lisher, 2013 [10] Saiab goatfish Indian Ocean off coast of Mozambique
Upeneus seychellensis Uiblein & Heemstra, 2011 Tailstripe goatfish Seychelles Bank in the western Indian Ocean
Upeneus stenopsis Uiblein & McGrouther, 2012 Narrowtail goatfish Western Pacific Ocean
Upeneus suahelicus Uiblein & Heemstra, 2010 Swahili goatfish Kenya to South Africa, Eritrea
Upeneus subvittatus Temminck & Schlegel, 1843 Deep-water goatfish Japan to Indonesia
Upeneus sulphureus G. Cuvier, 1829 Sulphur goatfish Indo-West Pacific
Upeneus sundaicus Bleeker, 1855 Ochrebanded goatfish Pakistan, India and Sri Lanka
Upeneus supravittatus Uiblein & Heemstra, 2010 Long-fin goatfish Sri Lanka, southern India
Upeneus taeniopterus G. Cuvier, 1829 Finstripe goatfish Indo-Pacific
Upeneus torres Uiblein & Gledhill, 2015 [11] Torres goatfish Australia, Vanuatu
Upeneus tragula J. Richardson, 1846 Freckled goatfish Indo-West Pacific
Upeneus vanuatu Uiblein & Causse, 2013 [12] Vanuatu goatfish Pacific Ocean, only known from Vanuatu
Upeneus vittatus Forsskål, 1775 Yellowstriped goatfish Indo-Pacific
Upeneus xanthogrammus C. H. Gilbert, 1892 Eastern Pacific


  1. ^ Cuvier, G. (1829): Le Règne Animal, distribué d'après son organisation, pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Edition 2. 2: i-xv + 1-406.
  2. ^ Uiblein, F. & Heemstra, P.C. (2010): A taxonomic review of the western Indian Ocean goatfishes of the genus Upeneus (Family Mullidae), with descriptions of four new species. Smithiana, Publications in Aquatic Biodiversity, Bulletin, 11: 35-71.
  3. ^ Lieske, E. & Myers, R. (1994): Collins Pocket Guide. Coral reef fishes. Indo-Pacific & Caribbean including the Red Sea. Harper Collins Publishers, 400 p.
  4. ^ a b Uiblein, F. & Gouws, G. (2014): A new goatfish species of the genus Upeneus (Mullidae) based on molecular and morphological screening and subsequent taxonomic analysis. Marine Biology Research, 10 (7): 655-681.
  5. ^ Yamashita, M., Golani, D. & Motomura, H. (2011). "A new species of Upeneus (Perciformes: Mullidae) from southern Japan". Zootaxa. 3107: 47–58.
  6. ^ Uiblein, F. & White, W.T. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Lombok, Indonesia and first verified record of U. asymmetricus for the Indian Ocean" (PDF). Zootaxa. 3980 (1): 51–66. doi:10.11646/zootaxa.3980.1.3.
  7. ^ "Upeneus moluccensis". CIESM Atlas of Exotic Fishes in the Mediterranean Sea. CIESM. Retrieved 15 January 2017.
  8. ^ Bos, A.R. (2014). "Upeneus nigromarginatus, a new species of goatfish (Perciformes: Mullidae) from the Philippines" (PDF). Raffles Bulletin of Zoology. 62: 745–753. Archived from the original (PDF) on 2014-12-13. Retrieved 2014-12-13.
  9. ^ "Upeneus pori". CIESM Atlas of Exotic Fishes in the Mediterranean Sea. CIESM. Retrieved 15 January 2017.
  10. ^ Uiblein, F. & Lisher, M. (2013): A new goatfish of the genus Upeneus (Mullidae) from Angoche, northern Mozambique. Zootaxa, 3717 (1): 85–95.
  11. ^ Uiblein F. & Gledhill D.C. (2015). "A new goatfish of the genus Upeneus (Mullidae) from Australia and Vanuatu, with inter- and intraspecific comparisons". Marine Biology Research. 11 (5): 475–491. doi:10.1080/17451000.2014.958088.
  12. ^ Uiblein, F. & Causse, R. (2013): A new deep-water goatfish of the genus Upeneus (Mullidae) from Vanuatu, South Pacific. Zootaxa, 3666 (3): 337–344.