The Refuge is one of only two (the other is Silvio O. Conte National Fish and Wildlife Refuge) that are located in parts of four states. As of 30 September 2007 the area per state was: Wisconsin: 89,637.54 acres (362.75 km²), Iowa: 51,147.78 acres (206.99 km²), Minnesota: 33,868.64 acres (137.06 km²), Illinois: 33,489.57 acres (135.53 km²). The following counties border on or have land within the Upper Mississippi River National Wildlife and Fish Refuge. In each state, the counties are listed from north to south. The lakes and rivers within the refuge area of each county are also listed.