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Trochonanina mozambicensis.png
A live individual of Trochonanina mozambicensis
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
(unranked): clade Heterobranchia

clade Euthyneura
clade Panpulmonata
clade Eupulmonata
clade Stylommatophora
clade Sigmurethra
clade limacoid clade

Superfamily: Helicarionoidea
Family: Urocyclidae
Simroth, 1889[1]

See text

Urocyclidae is a family of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]

Some species within this family make and use love darts before mating.


The Urocyclidae are distributed in the Afrotropical Region, including Madagascar, the Comores and the islands of the Gulf of Guinea.[3][4]


The family Urocyclidae consists of 2 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),[2] that follows classification of Urocyclinae by Van Goethem (1977):[5]

  • subfamily Urocyclinae Simroth, 1889
  • subfamily Sheldoniinae Connolly, 1925 (1912)[6] - synonyms: Peltatinae Godwin-Austen, 1912;[7] Trochonanininae Connolly, 1912;[8] Trochozonitinae Iredale, 1914;[9] Ledoulxiinae Pilsbry, 1919;[10] Gymnarionidae Von Mol, 1970;[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002


Genera within the family Urocyclidae include:

Subfamily Urocyclinae
tribe Urocyclini

Tribe Dendrolimacini

  • Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini[2]

Tribe Leptichnini

  • Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini[2]

Tribe Upembellini

  • Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini[2]

Subfamily Sheldoniinae



The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:[3]

 limacoid clade 



























  1. ^ Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. page 62.
  2. ^ a b c d e f g h i j k l m n o p Bouchet P.; Rocroi J.-P.; Frýda J.; Hausdorf B.; Ponder W.; Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology. Hackenheim, Germany: ConchBooks. 47 (1-2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997. 
  3. ^ a b Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
  4. ^ Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5. 
  5. ^ a b c d Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
  6. ^ Connolly (1925). Annals and Magazine of Natural History (9)15: 467.
  7. ^ Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. page 124.
  8. ^ Connolly (1912). Annals of the South African Museum 11(3): 101.
  9. ^ Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.
  10. ^ Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.
  11. ^ a b Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.
  12. ^ Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.
  13. ^ Heckert A. B. (2008). "Comments on the proposed conservation of Buettneria Case, 1922 (Amphibia) 1". Bulletin of Zoological Nomenclature 65: 310-313. HTM.
  14. ^ Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
  15. ^ de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.

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