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Temporal range: Wenlock–Westphalian
Purkynia lata (Fritsch).jpg
Purkynia lata
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Myriapoda
Class: Diplopoda
Subclass: Chilognatha
Order: †Zosterogrammida
Wilson, 2005
Family: †Zosterogrammidae
Wilson, 2005
  • Casiogrammus
  • Purkynia
  • Zosterogrammus

Zosterogrammidae is an extinct family of millipedes containing three genera, each with a single species. Fossils are known from the Czech Republic, Scotland, and the USA. Zosterogrammidae constitutes the sole family of the order Zosterogrammida.


Zosterogrammids have broad heads, 20 to 32 body segments, with a body tapering towards the head and rear. The dorsal plates of each segment are very broad and the rear portion of each is segment ornamented with lines. Size ranges from 26 millimetres (1.0 in) to 55 millimetres (2.2 in) in length. Superficially they are similar to the living Polyzoniida in body proportion.[1]

Taxonomy and distribution[edit]

Zosterogrammidae is placed in its own order, Zosterogrammida. Zosterogrammida is considered incertae sedis (uncertain placement) within the millipede subclass Chilognatha.[1]

Family †Zosterogrammidae Wilson, 2005
Casiogrammus ichthyeros Wilson, 2005 - Lanarkshire, Scotland; Wenlock (Silurian)
Purkynia lata Fritsch, 1899 - Nýřany, Czech Republic; Westphalian (Upper Carboniferous)
Zosterogrammus stichostethus Wilson, 2005 - Illinois, USA; Pennsylvanian (Upper Carboniferous)


  1. ^ a b Wilson, Heather M. (2005). "Zosterogrammida, a new order of millipedes from the middle Silurian of Scotland and the Upper Carboniferous of Euramerica". Palaeontology. 48 (5): 1101–1110. doi:10.1111/j.1475-4983.2005.00498.x.