Pyrgus

Grizzled skippers
	Grizzled skipper (Pyrgus malvae )
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Family:	Hesperiidae
Tribe:	Pyrgini
Genus:	Pyrgus; Hübner, [1819]
Synonyms
	Urbanus Hübner, [1806] (suppressed); Scelotrix Rambur, 1858; Bremeria Tutt, 1906; Teleomorpha Warren, 1926; Hemiteleomorapha Warren, 1926; Ateleomorpha Warren, 1926;

Pyrgus is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera Burnsius and Carterocephalus. They occur in the Holarctic with an additional group of species extending to the Neotropic.

In 2019, most of the species of Pyrgus found in the North, Central, or South America were moved to the genera Burnsius, Chirgus, and Heliopetes. The remaining Pyrgus species found in the New World are Pyrgus centaureae, ruralis, scriptura, and xanthus.^[1]^[2]

Species

These species belong to the genus Pyrgus:^[1]^[3]^[4]^[5]^[6]

Pyrgus accretus (Verity, 1925)
Pyrgus aladaghensis de Prins & van der Poorten, 1995 (aladag skipper)
Pyrgus alpinus Erschoff, 1874
Pyrgus alveus (Hübner, 1803) (large grizzled skipper)
Pyrgus andromedae (Wallengren, 1853) (alpine grizzled skipper)
Pyrgus armoricanus (Oberthür, 1910) (oberthür's grizzled skipper)
Pyrgus bieti (Oberthür, 1886)
Pyrgus bolkariensis de Prins & van der Poorten, 1995 (bolkar skipper)
Pyrgus cacaliae (Rambur, 1839) (dusky grizzled skipper)
Pyrgus carlinae (Rambur, 1839) (carline skipper)
Pyrgus carthami (Hübner, 1813) (safflower skipper)
Pyrgus centaureae (Rambur, 1842) (northern grizzled skipper)
Pyrgus centralitaliae (Verity, 1920)
Pyrgus chapmani Warren, 1926
Pyrgus cinarae (Rambur, 1839) (sandy grizzled skipper)
Pyrgus cirsii (Rambur, 1839) (cinquefoil skipper)
Pyrgus foulquieri (Oberthür, 1910) (foulquier's grizzled skipper)
Pyrgus iliensis (Reverdin, 1912)
Pyrgus jupei (Alberti, 1967) (caucasian skipper)
Pyrgus maculatus Bremer & Grey, 1853
Pyrgus malvae (Linnaeus, 1758) (grizzled skipper)
Pyrgus malvoides (Elwes & Edwards, 1897) (southern grizzled skipper)
Pyrgus melotis Duponchel, 1832 (aegean skipper)
Pyrgus nepalensis Higgins, 1984
Pyrgus onopordi (Rambur, 1839) (rosy grizzled skipper)
Pyrgus picenus Verity, 1920
Pyrgus pontica Reverdin, 1914
Pyrgus ruralis (Boisduval, 1852) (two-banded checkered skipper)
Pyrgus scriptura (Boisduval, 1852) (small checkered skipper)
Pyrgus serratulae (Rambur, 1839) (olive skipper)
Pyrgus sibirica (Reverdin, 1911)
Pyrgus sidae (Esper, 1784) (yellow-banded skipper)
Pyrgus speyeri (Staudinger, 1887)
Pyrgus warrenensis (Verity, 1928) (warren's skipper)
Pyrgus xanthus W. H. Edwards, 1878 (mountain checkered skipper)

References and external links

^ ^a ^b Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". Proceedings of the National Academy of Sciences. 116 (13): 6232–6237. Bibcode:2019PNAS..116.6232L. doi:10.1073/pnas.1821304116. PMC 6442542. PMID 30877254.
^ "Pyrgus genus Information". BugGuide.net. Retrieved 2023-03-30.
^ "Pyrgus". GBIF. Retrieved 2021-09-30.
^ Zhang, J; Cong, Q; Burns, JM; Grishin, NV (2022). "Checking the checkered taxonomy of Plötz's checkered skippers (Hesperiidae: Pyrgini)". Taxon Rep Int Lepid Surv. 10 (5). doi:10.5281/zenodo.6510282. PMC 9076161. PMID 35530660.
^ Pelham, Jonathan P. (2021). A Catalogue of the Butterflies of the United States and Canada (Report).
^ Zhang, Jing; Cong, Qian; Shen, Jinhui; Opler, Paul A.; et al. (2019). "Changes to North American butterfly names" (PDF). The Taxonomic Report of the International Lepidoptera Survey. 8 (2): 1–12. ISSN 2643-4806. PMC 8570546. PMID 34746798.

[Li2019-1] Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". Proceedings of the National Academy of Sciences. 116 (13): 6232–6237. Bibcode:2019PNAS..116.6232L. doi:10.1073/pnas.1821304116. PMC 6442542. PMID 30877254.

[bugguide-2] "Pyrgus genus Information". BugGuide.net. Retrieved 2023-03-30.

[gbif-3] "Pyrgus". GBIF. Retrieved 2021-09-30.

[Zhang2022-4] Zhang, J; Cong, Q; Burns, JM; Grishin, NV (2022). "Checking the checkered taxonomy of Plötz's checkered skippers (Hesperiidae: Pyrgini)". Taxon Rep Int Lepid Surv. 10 (5). doi:10.5281/zenodo.6510282. PMC 9076161. PMID 35530660.

[Pelham2021-5] Pelham, Jonathan P. (2021). A Catalogue of the Butterflies of the United States and Canada (Report).

[Zhang2019-6] Zhang, Jing; Cong, Qian; Shen, Jinhui; Opler, Paul A.; et al. (2019). "Changes to North American butterfly names" (PDF). The Taxonomic Report of the International Lepidoptera Survey. 8 (2): 1–12. ISSN 2643-4806. PMC 8570546. PMID 34746798.

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