Temporal range: Early Ordovician – Late Ordovician, 480–443Ma
|Life restoration of Arandaspis|
Ritchie and Gilbert-Tomlinson, 1977
Arandaspids (Subclass Arandaspida) are forming a group of very early, jawless prehistoric fish which lived during the Ordovician period. The group contains only one order, the Arandaspidiformes. The oldest known genus of this group is Sacabambaspis found in South America. Also, Arandaspids are one of the first craniotes, an unranked group of chordates.
The head armor of arandaspids is elongated, fusiform, with a rather flat dorsal shield, and a bulging ventral shield. In the anterior part of the dorsal shield are two closely set holes, which have been thought to be a paired pineal opening, but which are more likely the external openings of the endolymphatic ducts.
The eyes, surrounded by a sclerotic ring, are housed in a notch at the anterior end of the dorsal shield. The nostrils are nor clearly located, but may have been situated between the eyes. Ventrally, the ventral lip of the mouth is armed with long series of small oral plates which recall those of heterostracans.
The gill openings are probably numerous (more than 15) and minute. They opened between the diamond-shaped platelets which separate the dorsal from the ventral shield.
The body is covered with rod-shaped scales arranged in chevrons, and the tail is probably pad-shaped and diphycercal. The dermal bones of arandaspids consist of aspidine (acellular bone) and are ornamented with oakleaf-shaped tubercles which seem to contain no dentine. The sensory-lines were housed in narrow grooves between the tubercles.
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