Stratigraphic range: Cambrian
|Named for||Topographic feature (a summit) in Marathon County|
Blackberry Hill is a series of quarries and outcrops in Central Wisconsin that is notable for its large concentration of trace fossils in Cambrian rocks. One quarry in particular, located in Marathon County, also has the distinction of preserving some of the first land animals. The site is a prolific Cambrian Konservat-Lagerstätte. It includes three-dimensional casts of soft bodied and lightly scleritized invertebrates and a variety of exceptionally preserved types of trace fossils.
Age and stratigraphic placement
The strata at Blackberry Hill are known to belong to the Elk Mound Group; however, the lack of good stratigraphic markers (i.e., index fossils) in some Blackberry Hill localities, coupled with uncertainties about the age range of the Elk Mound Group itself, make it difficult to assign a precise age to these strata. Many researchers consider these rocks to be Late Cambrian, which is the age to which the Elk Mound Group was originally assigned;however, some recent authors now believe the Elk Mound Group and the fossils of Blackberry Hill could date back to the Middle Cambrian, based on certain fossils obtained from other areas.
Geological and environmental setting
Most of the strata are composed of well bedded quartz sandstone and orthoquartzite. They were deposited mainly on intertidal and supratidal sand flats along the southern shoreline of the supercontinent Laurentia. Ripple marks and numerous other sedimentary structures identical to those found on modern beaches abound on the strata surfaces. One of the most conspicuous features is extensive areas of specific structures not unlike those associated with modern biofilms and microbial mats. There is mounting evidence suggesting that the feeding potential of this presumed microbial material was one of the forces that lured the first animals out of the sea. It is also believed that the same material aided in the exceptional preservation of many of Blackberry Hill’s trace fossils.
Among the many paleontological discoveries thus far made at Blackberry Hill are the following:
- The first body fossils, and therefore the identity, of one of the first animals to walk on land.
- The only fossils that link those animals with their trackways. 
- The oldest body fossils of a phyllocarid (and perhaps the first crustaceans).
- Identification of the maker of Protichnites trackways after over 150 years of speculation.
- The largest concentrations of fossilized, stranded scyphozoans (i.e., jellyfish).
- What may be the first mating behavior in the fossil record.
The largest, most productive quarry is still in operation, thereby revealing fresh surfaces and the potential for new discoveries on a continuing basis.
Sedimentary structures associated with biofilms and microbial mats  are the only evidence of non-animal life at Blackberry Hill, as is the case elsewhere in this pre-embryophyte period in the history of Earth’s life on land. The animal life of Blackberry Hill was, however, represented by several kinds of macrofossils, all preserved as three-dimensional casts or impressions, including:
- Mosineia macnaughtoni– These large (up to 10+ cm in length) arthropods, apparently having paddle or oar-like appendages, along with another genus from similar-aged strata of Quebec, have been identified as the oldest euthycarcinoids in the fossil record and some of the first animals to walk on land. It is thus far known from only three exoskeletons preserved ventral-side-up, all lacking the heads and carapaces.
- Arenosicaris inflata– This smaller arthropod (up to approximately 5 cm in length), more common than Mosineia (over 40 specimens to date), is thought to be the oldest member of the Phyllocarida, and possibly the oldest crustacean in the fossil record. Disarticulated or splayed carapaces have been found in addition to essentially complete exoskeletons; however, the head is poorly known. Limbs, which are rarely fossilized in phyllocarids, are typically preserved in the A. inflata specimens.
- Scyphozoan medusae – These dome-shaped fossils are usually represented as body cavity infillings formed as the stranded animals pumped their bodies in an attempt to escape, and in so doing, ingested sand. Being composed mainly of water, scyphozoans rarely fossilize; however, they are found by the hundreds on some surfaces of one Blackberry Hill quarry.  The size of these fossils is also noteworthy; some specimens have achieved a diameter of 950 cm, making them the largest scyphozoan fossils on record. 
- Protichnites – These trackways (up to 8 cm wide at Blackberry Hill) are characterized by two parallel rows of paired footprints, often in sets, and a medial furrow or series of medial impressions, presumably from a dragging or touching tail. The maker of Protichnites was conjectural since 1852, when Sir Richard Owen, the pioneering British anatomist and paleontologist who coined the term “dinosauria,” first named and described them based on material from equivalent strata of Quebec. Fossils from Blackberry Hill named P. eremita,  found over 150 years later, solved the mystery by eventually revealing that at least some Protichnites were likely produced by the euthycarcinoid Mosineia macnaughtoni. P. eremita was originally interpreted to have been the product of an arthropod with the assistance of a mollusk shell that it used in hermit crab-style behavior.
- Diplichnites – These trackways at Blackberry Hill often resemble Protichnites, except they lack medial markings. They may have been made by the same animal(s) that produced Protichnites,  based on individual trackways having medial furrows that appear and disappear as the animal travelled. It has been suggested that some Diplichnites are undertracks, whereby the animal walked on the top layer of sand, but only the tips of its appendages reached the underlying layer. In these cases, the top layer might then exhibit Protichnites due to the tail that dragged on the surface, and the bottom layer would result in the furrow-less Diplichnites. It is possible that some Diplichnites were produced by trilobites or unknown arthropods, but no fossils of trilobites have been found thus far at Blackberry Hill.
- Climactichnites – These distinctive trace fossils, up to 14 cm in width and resembling tire tracks, are the most conspicuous trackways at Blackberry Hill, literally covering large surfaces of certain strata. Two ichnospecies are found: C. wilsoni, which are surface trackways that have lateral ridges; and C. youngi, which are sub-surface burrows that lack lateral ridges.  Their maker was likely a large, slug-like mollusk.  Occasionally the resting trace, Musculopodus, is found at one end of a C. wilsoni trackway.  It has been speculated that the animal may have remained burrowed in the sediment in the daytime to avoid desiccation from the sunlight,  and emerged at night to feed on the near-shore and on-shore microbial mats; however, there is no direct fossil evidence of grazing at Blackberry Hill, such as fossilized fecal pellets or strands.
- Other trace fossils – A variety of less conspicuous trace fossils is common at Blackberry Hill. The resting trace Rusophycus, bilobate linear traces similar to Cruziana, and ropelike traces are sometimes found in close association with Arenosicaris inflata and are believed to result mainly from the burrowing activity of that phyllocarid. 
- Collette and Hagadorn, 2010
- Hagadorn et al., 2002
- Schieber et al., 2007
- Getty and Hagadorn, 2008
- Ostrom, 1966
- Seilacher and Hagadorn, 2010
- Young and Hagadorn, 2010
- Collette et al., 2012
- MacNaughton et al., 2002
- Seilacher, 2007, page 28
- Seilacher, 2008
- Hagadorn et al., 2002
- Hagadorn and Belt, 2008, page 429
- Owen, 1852
- Hagadorn and Seilacher, 2009
- Goldring and Seilacher, 1971
- Hoxie, 2005
- Yochelson and Fedonkin, 1993
- Getty and Hagadorn, 2009
- Collette et al, 2010
- Collette, J. H., K. C. Gass, & J. W. Hagadorn (2012). "Protichnites eremita unshelled? Experimental model-based neoichnology and new evidence for a euthycarcinoid affinity for this ichnospecies". Journal of Paleontology 86 (3): 442–454. doi:10.1666/11-056.1.
- Collette, J. H. and J. W. Hagadorn (2010). "Three-dimensionally preserved arthropods from Cambrian Lagerstatten of Quebec and Wisconsin". Journal of Paleontology 84 (4): 646–667. doi:10.1666/09-075.1.
- Collette, J. H., J. W. Hagadorn, and M. A. LaCelle (2010). "Dead in their tracks: Cambrian arthropods and their traces from intertidal sandstones of Quebec and Wisconsin". PALAIOS 25 (8): 475–486. doi:10.2110/palo.2009.p09-134r.
- Getty, P. R., and J. W. Hagadorn (2008). "Reinterpretation of Climactichnites Logan 1860 to include subsurface burrows, and erection of Musculopodus for resting traces of the trailmaker". Journal of Paleontology 82 (6): 1161–1172. doi:10.1666/08-004.1.
- Getty, P. R., and J. W. Hagadorn (2009). "Palaeobiology of the Climactichnites tracemaker". Palaeontology 52 (4): 753–778. doi:10.1111/j.1475-4983.2009.00875.x.
- Goldring, R., and A. Seilacher (1971). "Limulid undertracks and their sedimentological implications". Neues Jarbuch fur Geologie und Palaontologie Abhandlungen 137: 422–442.
- Hagadorn, J. W. and E. S. Belt (2008). "Stranded in upstate New York: Cambrian scyphomedusae from the Potsdam Sandstone". PALAIOS 23 (7): 424–441. doi:10.2110/palo.2006.p06-104r.
- Hagadorn, J. W., R. H. Dott, and D. Damrow (2002). "Stranded on an Upper Cambrian Shoreline: Medusae from Central Wisconsin". Geology 30 (2): 147–150. doi:10.1130/0091-7613(2002)030<0147:SOALCS>2.0.CO;2.
- Hagadorn, J. W., and A. Seilacher (2009). "Hermit arthropods 500 million years ago?". Geology 37 (4): 295–298. doi:10.1130/G25181A.1.
- Hoxie, C. T. (2005). "Late Cambrian arthropod trackways in subaerially exposed environments: Incentives to simplify a problematic ichnogenus". Unpublished B.A. Thesis: 1–89.
- MacNaughton, R. B., J. M. Cole, R. W. Dalrymple, S. J. Braddy, D. E. G. Briggs, and T. D. Lukie (2002). "First steps on land:Arthropod trackways in Cambrian-Ordovician eolian sandstones, Canada". Geology 30 (5): 391–394. doi:10.1130/0091-7613(2002)030<0391:FSOLAT>2.0.CO;2.
- Ortega-Hernandez, J., Tremewan, J., and Braddy, S. J. (2010). "Euthycarcinoids". Geology Today 26 (5): 195–198. doi:10.1111/j.1365-2451.2010.00770.x.
- Ostrom, M. E. (1966). Cambrian Stratigraphy of Western Wisconsin. Wisconsin Geological and Natural History Survey.
- Owen, R. (1852). "Description of the impressions and footprints of the Protichnites from the Potsdam sandstone of Canada". Geological Society of London Quarterly Journal 8: 214–225. doi:10.1144/GSL.JGS.1852.008.01-02.26.
- Schieber, J.; Bose, P. K.; Eriksson, P. G.; Banerjee, S.; Sarkar, S.; Altermann, W.; Catuneau, O. (2007). Atlas of Microbial Mat Features Preserved within the Clastic Rock Record. Elsevier. pp. 53–71.
- Seilacher, A. (2007). Trace Fossil Analysis. New York, New York: Springer. p. 226.
- Seilacher, A. (2008). "Biomats, biofilms, and bioglue as preservational agents for arthropod trackways". Palaeogeography, Palaeoclimatology, Palaeoecology 270 (3–4): 252–257. doi:10.1016/j.palaeo.2008.07.011.
- Seilacher, A. and J. W. Hagadorn (2010). "Early molluscan evolution: Evidence from the trace fossil record". PALAIOS 25 (9): 565–575. doi:10.2110/palo.2009.p09-079r.
- Yochelson, E. L. and M. A. Fedonkin (1993). "Paleobiology of Climactichnites, an enigmatic Late Cambrian fossil". Smithsonian Contributions to Paleobiology 74: 1–74. doi:10.5479/si.00810266.74.1.
- Young, G. A. and J. W. Hagadorn (2010). "The fossil record of cnidarian medusa". Palaeoworld 19 (3–4): 212–221. doi:10.1016/j.palwor.2010.09.014.
- Provides a discussion and photographs of Blackberry Hill fossils, as one of Paleontology Portal’s “Famous Flora and Fauna” pages.