|Leaves and acorn|
Quercus chrysolepis, commonly termed Canyon Live Oak, or Golden Cup Oak, is a species of evergreen oak that is found in the southwestern part of North America, notably in the California Coast Ranges. This tree is often found near creeks and drainage swales growing in moist cool microhabitats. Its leaves are a glossy dark green on the upper surface with prominent spines; a further rapid identification arises from the leaves of Canyon live oak being geometrically flat. They are often sympatric with Quercus agrifolia and several other oak species. Fossil data supports a much wider distribution throughout the western United States during the early Holocene period.
Native Americans used the acorns of this species as a food staple, after leaching of the tannins; moreover, its roasted seed is a coffee substitute. After forest fires, Canyon live oak regenerates vigorously by basal sprouting, and the clonal diversity of this species has been shown to be high. Alternate common names for this taxon are Canyon Oak and Golden-Cup Oak.
Quercus chrysolepis is an evergreen tree with significant-sized spreading, horizontal branches, and a broad, rounded crown; it attains a height of six to thirty meters and is often found in a shrubby growth form. The trunk diameter can range from 30 to 100 centimeters. The elliptical to oblong leaves are 2.5 to 8.0 centimeters in length with widths of about half that dimension; moreover, leaves are short-pointed at the tip, but rounded or blunt at base. Although the leaves appear generally flat, they may have edge margins slightly turned under, typically with spiny teeth, particularly on young twigs. These leathery leaves are a glossy dark green above, with a nether surface a dull golden down, often becoming gray and nearly glabrous the second year.
Bark of the Canyon live oak is of a light gray coloration, and is rather smooth or sometimes scaly. Acorns occur solitarily or in pairs, exhibiting lengths of two to five centimeters; these fruits are variable in size and shape, but generally ovoid, turban-like with a shallow, thick cup of scales densely covered with yellowish hairs; the stalk is barely evident.
Distribution and habitat 
Quercus chrysolepis, Canyon live oak, is found in a variety of forest communities in the southwestern United States. It is found in 1.12 million acres (4,500 km²) of forest in California alone, including coast ranges and discontinuous occurrence on the eastern slopes of the southern and central Sierra Nevada. This tree is rare east of the Cascade-Sierra crest. Scattered, disjunct populations are found in the mountainous regions of southwest Nevada and in portions of western and central Arizona, and northwest Mexico.
Canyon live oak is tolerant of a variety of soil types, including very rocky or cobbly environments. It is hardy to cold temperatures down to - 11 °F, and will grow in neutral to moderately acidic soils with pH ranges of 4.5 to 7.5. An example of very rocky and serpentine soil tolerance is the species occurrence at the Cedars of Sonoma County, California. Canyon live oak grows at elevations of about 500 to 1500 meters in southwestern Oregon; in Northern California, from 100 to 1400 meters; and in Southern California, up to approximately 2700 meters. Quercus chrysolepis can be the dominant tree on steep canyon walls, especially in locations of shallow rocky soils. In areas of moderate to high rainfall, it occurs on south facing slopes, and in the hotter, drier parts of its distribution, on northerly slope faces.
Besides the prehistoric use of Canyon live oaks as a human food source, the acorns are consumed by a variety of wildlife as diverse as Acorn Woodpecker, California ground squirrel, dusky-footed wood rat, western harvest mouse and black-tailed deer. Surprisingly there seems little difference in food preference by wildlife among different oak species. Extensive hybridization of Quercus chrysolepis has been documented with several other sympatric oak species, probably to a greater extent than for any other Quercus species. The ability of Quercus chrysolepis to compete with other dominant trees within its range has been analyzed from the standpoint of leaf architecture and photosynthetic capability. The study results explain that, in low light environments, Q. chrysolepis out-competes species with superior leaf size and crown mass per unit volume by its greater photosynthetic efficiency and leaf lifespan.
Canyon live oak gives functional habitat for many fauna by providing perching, nesting, resting, or foraging sites for numerous species of birds, and shade and cover for diverse other mammals. Young Q. chrysolepis readily available browse. Canyon live oak woodlands serve as excellent mountain lion habitat because of the large population of deer frequenting these areas. Many species forage on Canyon live oak foliage including black-tailed jackrabbit, beaver, brush rabbit, red-backed vole, Sonoma chipmunk, cactus mouse, deer mouse, and porcupine. Pocket gophers often feed on the cambium of young canyon live oaks.
See also 
- California oak woodland
- California chaparral and woodlands
- Riparian woodland
- A.M. Montalvo, S.G. Conard, M.T. Conkle and P.D. Hodgskiss, Population structure, genetic diversity, and clone formation in Quercus chrysolepis (Fagaceae), American Journal of Botany, Vol 84, 1553, (1997)
- Bruce M. Pavlik, Pamela C. Muick, Sharon Johnson and Marjorie Popper, Oaks of California, Cachuma Press (1992) ISBN 0-9628505-1-9
- C.Michael Hogan. 2010. Leather oak. eds. Mark McGinley & C.J.Cleveland. Encyclopedia of Earth. National Council for Science and the Environment. Washington DC
- McDonald, Philip M., and Edward E. Littrell, 1976. The bigcone Douglas-fir-canyon live oak community in southern California, Madroño 23:310-320
- Peter G. Kennedy, Post-dispersal seed predation varies by habitat not acorn size for Quercus chrysolepis (Fagaceae) and Lithocarpus Densiflora (Fagaceae) in central coastal California, Madrono: Vol. 52, No. 1, pp. 30–34
- Kevin C. Brown, Biodiversity of Oak (Quercus) Species in California and Adjacent Regions, Fifth Symposium on Oak Woodlands, San Diego, Ca., Oct. 22-25, 2001
- J. C. Hunter, Correspondence of environmental tolerances with leaf and branch attributes for six co-occurring species of broadleaf evergreen trees in northern California, Journal: Trees - Structure and Function, Volume 11, Number 3, Pages 169-175, January, 1997; Springer, Berlin / Heidelberg, ISSN 0931-1890
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