Helius (fly)

Helius
Helius flavus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Suborder:	Nematocera
Infraorder:	Tipulomorpha
Superfamily:	Tipuloidea
Family:	Limoniidae
Subfamily:	Limoniinae
Genus:	Helius Le Peletier & Serville, 1828
Type species
Limnobia longirostris Meigen, 1818
Subgenera
Eurhamphidia Alexander, 1915 Helius Lepeletier & Serville, 1828 Idiohelius Alexander, 1948 Mammuthonasus Theischinger, 1994 Prohelius Alexander, 1975 Rhamphidina Alexander, 1920 Rhamphidioides Alexander, 1920 Rhampholimnobia Alexander, 1915 Rhyncholimonia Alexander, 1964
Synonyms
Megarhina Le Peletier & Serville, 1828 Leptorhina Stephens, 1829 Rhamphidia Meigen, 1830

Helius is a genus of crane fly in the family Limoniidae, distributed worldwide with most species in Australia and East Asia.

Species

• Subgenus Eurhamphidia Alexander, 1915

• H. abnormalis (Brunetti, 1918)
• H. ata Alexander, 1931
• H. atroapicalis Alexander, 1978
• H. auranticolor Alexander, 1936
• H. connectus Edwards, 1928
• H. diacanthus Alexander, 1931
• H. fuscofemoratus Alexander, 1931
• H. glabristylatus Alexander, 1931
• H. indivisus Alexander, 1931
• H. inelegans (Alexander, 1928)
• H. invenustipes (Alexander, 1930)
• H. melanosoma Alexander, 1936
• H. mimicans Edwards, 1933
• H. mirus Edwards, 1926
• H. monticola (Edwards, 1919)
• H. mouensis Alexander, 1948
• H. nigrofemoratus (Alexander, 1930)
• H. niveitarsis (Skuse, 1890)
• H. pallens Edwards, 1933
• H. perelegans Alexander, 1930
• H. perlongatus Alexander, 1978
• H. scabiosus Alexander, 1932
• H. vitiensis Alexander, 1956

• Subgenus Helius Lepeletier & Serville, 1828

• H. acanthostyla Alexander, 1944
• H. aciferus Alexander, 1937
• H. aka Alexander, 1972
• H. albitarsis (Osten Sacken, 1888)
• H. albogeniculatus Alexander, 1936
• H. amplus Edwards, 1933
• H. anaemicus Alexander, 1932
• H. anamalaiensis Alexander, 1967
• H. angustalbus Alexander, 1953
• H. aphrophilus Alexander, 1948
• H. apicalis (Alexander, 1915)
• H. apoensis Alexander, 1931
• H. apophysalis Alexander, 1967
• H. araucariae Alexander, 1945
• H. arcuarius Alexander, 1929
• H. argyrosternus Alexander, 1930
• H. arunachalus Alexander, 1975
• H. attenuatus Alexander, 1929
• H. barbatus Edwards, 1921
• H. bicolor Edwards, 1933
• H. bifurcus Alexander, 1956
• H. bitergatus Alexander, 1950
• H. boops Alexander, 1942
• H. brachyphallus Alexander, 1956
• H. brevioricornis (Alexander, 1920)
• H. brevisector Alexander, 1956
• H. brunneus Byers, 1981
• H. cacoxenus (Alexander, 1920)
• H. calviensis Edwards, 1928
• H. capensis (Alexander, 1917)
• H. capniopterus Alexander, 1945
• H. catreus Alexander, 1967
• H. cavernicolus Alexander, 1961
• H. chikurinensis Alexander, 1930
• H. chintoo Theischinger, 1994
• H. communis (Skuse, 1890)
• H. comoreanus Alexander, 1959
• H. compactus Alexander, 1967
• H. copiosus Alexander, 1935
• H. corniger Savchenko, 1983
• H. costofimbriatus Alexander, 1930
• H. costosetosus Alexander, 1932
• H. creper Alexander, 1926
• H. ctenonycha Alexander, 1938
• H. dafla Alexander, 1972
• H. darlingtonae Welch & Gelhaus, 1994
• H. destitutus Alexander, 1971
• H. devinctus Alexander, 1932
• H. distinervis Alexander, 1940
• H. dolichorhynchus Edwards, 1933
• H. eremnophallus Alexander, 1978
• H. euryphallus Alexander, 1960
• H. fasciventris Edwards, 1926
• H. ferruginosus (Brunetti, 1912)
• H. flavidibasis Alexander, 1975
• H. flavipes (Macquart, 1855)
• H. flavitarsis (Alexander, 1920)
• H. flavus (Walker, 1856)
• H. fragosus Alexander, 1937
• H. franckianus Alexander, 1940
• H. fratellus (Brunetti, 1918)
• H. fulvithorax (Skuse, 1890)
• H. fumicosta Edwards, 1928
• H. fuscoangustus Alexander, 1967
• H. garcianus Alexander, 1972
• H. gibbifer Savchenko, 1981
• H. gorokanus Alexander, 1962
• H. gracillimus Alexander, 1938
• H. graphipterus Alexander, 1954
• H. haemorrhoidalis Alexander, 1937
• H. harrisi (Alexander, 1923)
• H. hatschbachi Alexander, 1954
• H. hispanicus Lackschewitz, 1928
• H. hova Alexander, 1955
• H. impensus Alexander, 1967
• H. imperfectus (Alexander, 1920)
• H. inconspicuus (Brunetti, 1912)
• H. ineptus Alexander, 1938
• H. infirmus Alexander, 1932
• H. invariegatus Alexander, 1938
• H. iris (Alexander, 1920)
• H. kambangani (de Meijere, 1913)
• H. kambanganoides Alexander, 1931
• H. larotypa Alexander, 1926
• H. lectus Alexander, 1936
• H. leucoplaca Alexander, 1936
• H. lienpingensis Alexander, 1945
• H. liguliferus Alexander, 1978
• H. liliputanus Alexander, 1929
• H. lobuliferus Alexander, 1938
• H. longinervis Edwards, 1928
• H. longirostris (Meigen, 1818)
• H. luachimoensis Alexander, 1963
• H. luniger (Riedel, 1914)
• H. mainensis (Alexander, 1916)
• H. malagasicus Alexander, 1955
• H. manueli Alexander, 1972
• H. medleri Alexander, 1976
• H. melanophallus Alexander, 1938
• H. mesorhynchus Alexander, 1933
• H. micracanthus Alexander, 1945
• H. mirabilis (Alexander, 1914)
• H. mirandus (Alexander, 1921)
• H. morosus (Alexander, 1920)
• H. multivolus Alexander, 1945
• H. murreensis Alexander, 1960
• H. myersiellus Alexander, 1935
• H. nawaianus Alexander, 1929
• H. neocaledonicus Alexander, 1945
• H. nigricapella Alexander, 1938
• H. nigriceps (Edwards, 1916)
• H. nipponensis (Alexander, 1913)
• H. numenius Alexander, 1967
• H. obliteratus (Alexander, 1920)
• H. obsoletus (Alexander, 1920)
• H. oxystylus Alexander, 1967
• H. pallidipes Alexander, 1926
• H. pallidissimus Alexander, 1930
• H. pallirostris Edwards, 1921
• H. panamensis Alexander, 1934
• H. paramorosus Alexander, 1949
• H. parvidens Alexander, 1944
• H. patens Edwards, 1933
• H. pauperculus (Alexander, 1921)
• H. pavoninus Alexander, 1938
• H. perflavens Alexander, 1964
• H. perpallidus Alexander, 1942
• H. pervenustus Alexander, 1953
• H. phasmatis Alexander, 1945
• H. pictus Edwards, 1926
• H. plebeius Alexander, 1946
• H. pluto Alexander, 1932
• H. polionotus Alexander, 1938
• H. procerus Alexander, 1931
• H. productellus Alexander, 1944
• H. protumidus Alexander, 1978
• H. quadrifidus Alexander, 1926
• H. quadrivena Alexander, 1971
• H. rectispinus Alexander, 1947
• H. rectus Alexander, 1945
• H. regius Alexander, 1944
• H. rostratus Alexander, 1956
• H. rubicundus (Alexander, 1922)
• H. rufescens (Edwards, 1916)
• H. rufithorax Alexander, 1928
• H. sanguinolentus (Alexander, 1921)
• H. schildi Alexander, 1945
• H. selectus Alexander, 1936
• H. serenus Alexander, 1967
• H. setigerus Alexander, 1980
• H. sigillatus Alexander, 1964
• H. stenorhynchus Alexander, 1954
• H. stolidus Alexander, 1948
• H. subanaemicus Alexander, 1964
• H. subarcuarius Alexander, 1934
• H. subfasciatus Alexander, 1924
• H. submorosus (Alexander, 1921)
• H. subobsoletus (Alexander, 1921)
• H. subpauperculus Alexander, 1975
• H. tantalus Alexander, 1967
• H. tanyrhinus Alexander, 1964
• H. taos Alexander, 1967
• H. tenuirostris Alexander, 1924
• H. tenuistylus Alexander, 1929
• H. tetracradus Alexander, 1967
• H. tienmuanus Alexander, 1940
• H. tonaleah Theischinger, 1994
• H. trianguliferus Alexander, 1931
• H. tshinta Theischinger, 1994
• H. unicolor (Brunetti, 1912)
• H. uniformis (Alexander, 1914)
• H. venustus (Skuse, 1890)
• H. verticillatus Alexander, 1967
• H. yindi Theischinger, 1994

• Subgenus Idiohelius Alexander, 1948

• H. melanolithus Alexander, 1953
• H. mirificus Alexander, 1953
• H. pentaneura Alexander, 1948

• Subgenus Mammuthonasus Theischinger, 1994

• H. allunga Theischinger, 1994

• Subgenus Prohelius Alexander, 1975

• H. dugaldi Alexander, 1945
• H. edwardsianus Alexander, 1956
• H. fulani Alexander, 1975

• Subgenus Rhamphidina Alexander, 1920

• H. camerounensis (Alexander, 1920)

• Subgenus Rhamphidioides Alexander, 1920

• H. alluaudi (Riedel, 1914)
• H. venustissimus (Alexander, 1920)

• Subgenus Rhampholimnobia Alexander, 1915

• H. bigeminatus Alexander, 1962
• H. brevinasus Alexander, 1934
• H. deentoo Theischinger, 1994
• H. diffusus Alexander, 1941
• H. fenestratus Alexander, 1973
• H. gracilirostris Alexander, 1962
• H. guttulinus Alexander, 1960
• H. japenensis Alexander, 1948
• H. matilei Hynes, 1993
• H. mesolineatus Alexander, 1962
• H. nimbus Alexander, 1941
• H. papuanus Alexander, 1934
• H. reticularis (Alexander, 1915)
• H. simulator Alexander, 1962
• H. spinteris Alexander, 1960
• H. subreticulatus Alexander, 1962

• Subgenus Rhyncholimonia Alexander, 1964

• H. dicroneurus Alexander, 1964

References

• "Catalogue of the Craneflies of the World (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae)". Naturalis Biodiversity Center.