Scarabaeinae

Scarabaeinae
Copris lunaris (Coprini) ♀
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Superfamily:	Scarabaeoidea
Family:	Scarabaeidae
Subfamily:	Scarabaeinae Latreille, 1802
Tribes
About one dozen; see text
Synonyms
Coprinae

The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.

Scarabaeinae tribes, with some selected genera and species, are:

• Canthonini
  • Anachalcos Hope, 1837
  • Circellium Latreille, 1825
  • Dicranocara Frolov and Scholtz, 2003
  • Versicorpus Deschodt Davis and Scholtz, 2011
  • Deltochilum
    • Deltochilum valgum
• Coprini
  • Copris Geoffroy, 1762
  • Heliocopris
• Dichotomiini
  • Dichotomius
• Eucraniini
• Eurysterini
  • Eurysternus
• Gymnopleurini (sometimes included in Scarabaeini)
  • Gymnopleurus Illiger, 1803
• Oniticellini
  • Euoniticellus Janssens, 1953
  • Oniticellus Serville, 1825
• Onitini
  • Bubas Mulsant, 1842
  • Cheironitis Lansberge, 1875
  • Onitis Fabricius, 1798
• Onthophagini
• Phanaeini MacLeay, 1819
  • Oxysternon
  • Phanaeus
    • Phanaeus changdiazi
    • Phanaeus vindex
• Scarabaeini
  • Scarabaeus
• Sisyphini (sometimes included in Scarabaeini)
  • Neosisyphus
  • Sisyphus
• Unknown tribe / Incertae sedis
  • Anisocanthon
  • Anomiopus

Coprini, Oniticellini, Onitini, Onthophagini and Phanaeini are sometimes separated as subfamily Coprinae.

• 
  Cheironitis scabrosus
• 
  Chironitis indicus
• 
  Copris elphenor female
• 
  Copris elphenor male

External links

• Scarabaeinae Research Network
• photos of species from each tribe
• African Dung Beetles Online