The dentition of Yacarerani, like many other notosuchians, is heterodont, with different tooth morphologies in different parts of the jaws. Two teeth in the lower jaw project forward from the tip, resembling the incisors of a rabbit. Other teeth, situated posteriorly, are cusped and adapted for grinding or chopping food such as tubers or small arthropods. In the lower jaw, the dentary tooth rows merge posterior to the anterior dentary teeth. The same is true for the upper tooth rows, which merge on the palate.
Yacarerani presents similar teeth morphology to Adamantinasuchus navae, a crocodylomorph from Adamantina Formation, Bauru Group, Brazil.
Below is a cladogram from Novas et al. (2009) showing the phylogenetic position of Yacarerani: