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{{Short description|Family of amphibians}}
{{automatic Taxobox
{{Automatic taxobox
| name = Ceratophryines
| name = Ceratophryidae
| fossil_range = [[Late Cretaceous]] – [[Holocene]], {{Fossil range|70|0}}
| taxon = Ceratophryidae
| fossil_range = {{fossilrange|Miocene|Recent}}
| image = Argentine Horned Frog (Ceratophrys ornata)1.JPG
| image = Argentine Horned Frog (Ceratophrys ornata)1.JPG
| image_caption = ''[[Ceratophrys ornata]]''
| image_caption = ''[[Ceratophrys ornata]]''
| authority = [[Johann Jakob von Tschudi|Tschudi]], 1838
| authority = [[Johann Jakob von Tschudi|Tschudi]], 1838
| subdivision_ranks = [[Genera]]
| subdivision_ranks = [[Genera]]
| subdivision =
| subdivision = 3 extant genera, see text.
See text.
}}
}}
The '''Ceratophryidae''', also known as '''common horned frogs''', are a family of [[frog]]s found in [[South America]]. It is a relatively small family with three [[Extant taxon|extant]] genera and 12 species.<ref name=frost>{{cite web |url=http://research.amnh.org/vz/herpetology/amphibia/Amphibia/Anura/Ceratophryidae |title=Ceratophryidae Tschudi, 1838 |author=Frost, Darrel R. |year=2014 |work=Amphibian Species of the World: an Online Reference. Version 6.0 |publisher=American Museum of Natural History |accessdate=18 December 2014}}</ref><ref name=AmphibiaWeb>{{cite web |url=http://www.amphibiaweb.org/lists/Ceratophryidae.shtml |title=Ceratophryidae |year=2014 |work=AmphibiaWeb: Information on amphibian biology and conservation. [web application] |publisher=Berkeley, California: AmphibiaWeb |accessdate=13 December 2014 |archive-url=https://web.archive.org/web/20141209121203/http://amphibiaweb.org/lists/Ceratophryidae.shtml |archive-date=9 December 2014 |url-status=dead }}</ref><ref name=Blackburn2011>{{cite journal |author1=Blackburn, D.C. |author2=Wake, D.B. |name-list-style=amp | title=Class Amphibia Gray, 1825. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness | journal=Zootaxa| volume=3148| year=2011| pages=39–55|doi=10.11646/zootaxa.3148.1.8 | url=http://mapress.com/zootaxa/2011/f/zt03148p055.pdf}}</ref> Despite the common name, not all species in the family have the horn-like projections at the eyes.{{Citation needed|date=December 2014}} They have a relatively large head with big mouth, and they are ambush predators able to consume large prey, including lizards, other frogs, and small mammals. They inhabit arid areas and are seasonal breeders, depositing many small eggs in aquatic habitats. [[Tadpole]]s are free-living and carnivorous (''[[Ceratophrys]]'' and ''[[Lepidobatrachus]]'') or grazers (''[[Chacophrys]]'').<ref name="VittCaldwell">{{cite book|title=Herpetology: An Introductory Biology of Amphibians and Reptiles |edition=4th |first1=Laurie J.|last1= Vitt|first2=Janalee P. |last2=Caldwell |name-list-style=amp |publisher=Academic Press|year=2014|pages=496–497}}</ref>


Some species (especially from the genera ''Ceratophrys'' and ''Lepidobatrachus'') are popular in [[herpetoculture]].{{Citation needed|date=December 2014}}
'''Ceratophryinae''', known as '''common horned frogs''', is a [[subfamily]] of [[Leptodactylidae|leptodactylid]] [[frog]]s found in [[South America]]. Fossils of the giant ''[[Beelzebufo]]'' from [[Cretaceous]] [[Madagascar]] suggest the subfamily may have once ranged throughout the prehistoric [[supercontinent]] of [[Gondwana]].<ref name="nsf">{{cite web |url= http://www.nsf.gov/news/news_summ.jsp?cntn_id=111119&org=NSF&from=news |title= Scientists Discover 'Giant Fossil Frog from Hell' |publisher= [[National Science Foundation]] |last= Dybas |first= Cheryl |work= Press Release 08-025 |date= 2008-02-18 |accessdate= 2008-02-19 }}</ref> Despite the common name, not all species in the subfamily have the horn-like projections at the eyes. All are [[carnivorous]], [[Terrestrial animal|terrestrial]] or semi[[Aquatic animal|aquatic]], and notably thickset with proportionally large heads and mouths. Some species (especially from the genera ''Ceratophrys'' and ''Lepidobatrachus'') are popular in [[herpetoculture]].


The oldest fossils of the family are known from the [[Miocene]] epoch. The fossil giant frog ''[[Beelzebufo]]'' from the [[Cretaceous|Late Cretaceous]] of [[Madagascar]] was formerly considered to belong to this family, but is now excluded, but is possibly closely related, alongside ''[[Baurubatrachus]]'' from the Late Cretaceous of Brazil. ''[[Wawelia (frog)|Wawelia]]'' from the Miocene of Argentina is no longer considered closely related.<ref name=":1">{{Cite journal |last1=Barcelos |first1=Lucas Almeida |last2=Almeida-Silva |first2=Diego |last3=Santos |first3=Charles Morphy D. |last4=Verdade |first4=Vanessa Kruth |date=2021-10-18 |title=Phylogenetic analysis of Ceratophryidae (Anura: Hyloidea) including extant and extinct species |url=https://www.tandfonline.com/doi/full/10.1080/14772019.2022.2050824 |journal=Journal of Systematic Palaeontology |language=en |volume=19 |issue=20 |pages=1449–1466 |doi=10.1080/14772019.2022.2050824 |s2cid=248653602 |issn=1477-2019}}</ref>

== Taxonomy ==
Placement of this clade has varied considerably over time, having been a [[subfamily]] within the [[Leptodactylidae]] for a long while. Later on, it has been raised to family level, either broadly defined, including the [[Telmatobiidae]] and [[Batrachylidae]] (as subfamilies Telmatobiinae and Batrachylinae, respectively<ref name=VittCaldwell/>), or as now is commonly accepted, as a smaller family with three genera.<ref name=frost/><ref name=AmphibiaWeb/><ref name=Blackburn2011/>
==Genera==
==Genera==
The [[Extant taxon|extant]] genera are:<ref name=frost/><ref name=AmphibiaWeb/>
* †''[[Beelzebufo]]'' <small>Evans, Jones, & Krause, 2008</small> (fossil)
* ''[[Ceratophrys]]'' <small>Wied-Neuwied, 1824</small>
* ''[[Ceratophrys]]'' <small>Wied-Neuwied, 1824</small> (8 species)
* ''[[Chacophrys]]'' <small>Reig & Limeses, 1963</small>
* ''[[Chacophrys]]'' <small>Reig & Limeses, 1963</small> ([[monotypic]]: ''[[Chacophrys pierottii]]'' <small>(Vellard, 1948)</small>)
** ''[[Chacophrys pierottii]]'' <small>(Vellard, 1948)</small>
* ''[[Lepidobatrachus]]'' <small>Budgett, 1899</small> (3 species)

* ''[[Lepidobatrachus]]'' <small>Budgett, 1899</small>
In addition, a number of fossil taxa have been considered to be closely related, including:<ref name=":1" />
* ''[[Macrogenioglottus]]'' <small>Carvalho, 1946</small>
** ''[[Macrogenioglottus alipioi]]'' <small>Carvalho, 1946</small>
* †''[[Beelzebufo]]'' <small>Evans, Jones, & Krause, 2008</small>
* ''[[Odontophrynus]]'' <small>Reinhardt & Lütken, 1862</small>
* †''[[Baurubatrachus]]'' <small>Báez and Perí, 1990</small>
* ''[[Proceratophrys]]'' <small>Miranda-Ribeiro, 1920</small>


==References==
==References==
{{reflist}}
{{Reflist|2}}
*http://research.amnh.org/herpetology/amphibia/names.php?taxon=&family=&subfamily=Ceratophryinae&genus=&commname=&authority=&year=&geo=0&dist=&comment=


{{Taxonbar|from=Q55461}}
⚫
[[Category:Leptodactylidae]]
⚫
[[Category:Ceratophryinae|*]]


⚫
[[Category:Ceratophryidae| ]]
{{Leptodactylidae-stub}}
⚫
[[Category:Amphibian families]]
[[Category:Turonian first appearances]]
[[Category:Taxa named by Johann Jakob von Tschudi]]
[[Category:Extant Turonian first appearances]]

Revision as of 07:10, 3 January 2023

Ceratophryidae
Temporal range: Miocene–Recent
Ceratophrys ornata
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Amphibia
Order: Anura
Superfamily: Hyloidea
Family: Ceratophryidae
Tschudi, 1838
Genera

3 extant genera, see text.

The Ceratophryidae, also known as common horned frogs, are a family of frogs found in South America. It is a relatively small family with three extant genera and 12 species.[1][2][3] Despite the common name, not all species in the family have the horn-like projections at the eyes.[citation needed] They have a relatively large head with big mouth, and they are ambush predators able to consume large prey, including lizards, other frogs, and small mammals. They inhabit arid areas and are seasonal breeders, depositing many small eggs in aquatic habitats. Tadpoles are free-living and carnivorous (Ceratophrys and Lepidobatrachus) or grazers (Chacophrys).[4]

Some species (especially from the genera Ceratophrys and Lepidobatrachus) are popular in herpetoculture.[citation needed]

The oldest fossils of the family are known from the Miocene epoch. The fossil giant frog Beelzebufo from the Late Cretaceous of Madagascar was formerly considered to belong to this family, but is now excluded, but is possibly closely related, alongside Baurubatrachus from the Late Cretaceous of Brazil. Wawelia from the Miocene of Argentina is no longer considered closely related.[5]

Taxonomy

Placement of this clade has varied considerably over time, having been a subfamily within the Leptodactylidae for a long while. Later on, it has been raised to family level, either broadly defined, including the Telmatobiidae and Batrachylidae (as subfamilies Telmatobiinae and Batrachylinae, respectively[4]), or as now is commonly accepted, as a smaller family with three genera.[1][2][3]

Genera

The extant genera are:[1][2]

In addition, a number of fossil taxa have been considered to be closely related, including:[5]

References

  1. ^ a b c Frost, Darrel R. (2014). "Ceratophryidae Tschudi, 1838". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 18 December 2014.
  2. ^ a b c "Ceratophryidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2014. Archived from the original on 9 December 2014. Retrieved 13 December 2014.
  3. ^ a b Blackburn, D.C. & Wake, D.B. (2011). "Class Amphibia Gray, 1825. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness" (PDF). Zootaxa. 3148: 39–55. doi:10.11646/zootaxa.3148.1.8.
  4. ^ a b Vitt, Laurie J. & Caldwell, Janalee P. (2014). Herpetology: An Introductory Biology of Amphibians and Reptiles (4th ed.). Academic Press. pp. 496–497.
  5. ^ a b Barcelos, Lucas Almeida; Almeida-Silva, Diego; Santos, Charles Morphy D.; Verdade, Vanessa Kruth (2021-10-18). "Phylogenetic analysis of Ceratophryidae (Anura: Hyloidea) including extant and extinct species". Journal of Systematic Palaeontology. 19 (20): 1449–1466. doi:10.1080/14772019.2022.2050824. ISSN 1477-2019. S2CID 248653602.
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