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The [[Type (biology)|type species]] (and only known species of ''Utahraptor''), ''Utahraptor ostrommaysorum'', was named by Kirkland, Gaston and Burge in 1993 for the [[United States|American]] [[paleontologist]] [[John Ostrom]], from Yale University's [[Peabody Museum of Natural History]], and Chris Mays, of [[Dinamation]] International. Originally in the [[specific name (zoology)|specific name]] the singular [[genitive]] ''ostrommaysi'' was used but in 2000 this was emended by [[George Olshevsky]] to the plural.<ref>Olshevsky, G., 2000, ''An annotated checklist of dinosaur species by continent. Mesozoic Meanderings'' '''3''': 1-157</ref> Earlier, it had been intended to name the species "Utahraptor spielbergi" after [[film director]] [[Steven Spielberg]], in exchange for funding palaeontological research, but no agreement could be reached on the amount of financial assistance.<ref>Brooke Adams, 1993, "Director Loses Utahraptor Name Game", ''Deseret News'', 15 June 1993</ref>
The [[Type (biology)|type species]] (and only known species of ''Utahraptor''), ''Utahraptor ostrommaysorum'', was named by Kirkland, Gaston and Burge in 1993 for the [[United States|American]] [[paleontologist]] [[John Ostrom]], from Yale University's [[Peabody Museum of Natural History]], and Chris Mays, of [[Dinamation]] International. Originally in the [[specific name (zoology)|specific name]] the singular [[genitive]] ''ostrommaysi'' was used but in 2000 this was emended by [[George Olshevsky]] to the plural.<ref>Olshevsky, G., 2000, ''An annotated checklist of dinosaur species by continent. Mesozoic Meanderings'' '''3''': 1-157</ref> Earlier, it had been intended to name the species "Utahraptor spielbergi" after [[film director]] [[Steven Spielberg]], in exchange for funding palaeontological research, but no agreement could be reached on the amount of financial assistance.<ref>Brooke Adams, 1993, "Director Loses Utahraptor Name Game", ''Deseret News'', 15 June 1993</ref>

==In popular culture==
A speculative reconstruction of ''Utahraptor'' lifestyles and behavior was presented in the 1999 [[BBC]] television series ''[[Walking with Dinosaurs]]''.<ref name=WWDmedia>"Unnatural history? Deconstructing the ''Walking with Dinosaurs'' phenomenon." ''Media Culture & Society'', '''25'''(3): 315-332. {{doi|10.1177/0163443703025003002}}</ref> The program portrayed ''Utahraptor'' with several anatomical inaccuracies including [[Pronation|pronated]] hands and featherless skin, and depicted them living in Europe when the only fossils of ''Utahraptor'' have been found in western North America.<ref name=WWDnathist>Haines, T. (2000). ''Walking with Dinosaurs: A Natural History''. London: BBC Publishing.</ref>


==References==
==References==

Revision as of 23:03, 10 February 2014

Utahraptor
Temporal range: Early Cretaceous, 126 Ma
Cast of foot bones
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Dromaeosauridae
Clade: Eudromaeosauria
Genus: Utahraptor
Kirkland, Gaston & Burge, 1993
Species:
U. ostrommaysorum
Binomial name
Utahraptor ostrommaysorum
Kirkland, Gaston & Burge, 1993

Utahraptor (meaning "Utah's predator"[1]) is a genus of theropod dinosaurs, including the largest known members of the family Dromaeosauridae. Fossil specimens date to the upper Barremian stage of the early Cretaceous period (in rock strata dated to 126 ± 2.5 million years ago).[2] It contains a single species, Utahraptor ostrommaysorum.

Description

Size of the largest individual (orange) of Utahraptor compared to a human

The holotype specimen of Utahraptor is fragmentary, consisting of skull fragments, a tibia, claws and some caudal (tail) vertebrae. These few elements suggest an animal about twice the length of Deinonychus.[1] Like other dromaeosaurids, Utahraptor had large curved claws on their second toes. One claw specimen is preserved at 22 centimetres (8.7 in) in length and is thought to reach 24 centimetres (9.4 in) restored.

The largest described U. ostrommaysorum specimens are estimated to have reached up to 7 m (23 ft) long and somewhat less than 500 kg (1,100 lb) in weight, comparable to a polar bear in size.[1][3]

It is thought that Utahraptor may be closely related to the smaller Dromaeosaurus and the giant Mongolian dromaeosaurid genus Achillobator.[1][4]

Although feathers have never been found in association with Utahraptor specimens, there is strong phylogenetic evidence suggesting that all dromaeosaurids possessed them. This evidence comes from phylogenetic bracketing, which allows paleontologists to infer traits that exist in a clade based on the existence of that trait in a more basal form. The genus Microraptor is one of the oldest known dromaeosaurids, and is phylogenetically more primitive than Utahraptor.[5] Since Microraptor and other dromaeosaurids possessed feathers, it is reasonable to assume that this trait was present in all of Dromaeosauridae. Feathers were very unlikely to have evolved more than once, so assuming that any given dromaeosaurid, such as Utahraptor, lacked feathers would require positive evidence that they did not have them.[6] So far, there is nothing to suggest that feathers were lost in larger, more derived species of dromaeosaurs.[7]

In a 2001 study conducted by Bruce Rothschild and other paleontologists, 2 foot bones referred to Utahraptor were examined for signs of stress fracture, but none were found.[8]

Discovery

Life restoration based on known specimens

The first specimens of Utahraptor were found in 1975 by Jim Jensen in the Dalton Wells Quarry in east-central Utah, near the town of Moab, but did not receive much attention. After a find of a large foot-claw by Carl Limoni in October 1991 James Kirkland, Robert Gaston, and Donald Burge uncovered further remains of Utahraptor in 1991 in the Gaston Quarry in Grand County, Utah, within the Yellow Cat and Poison Strip members of the Cedar Mountain Formation.[1] Radiometric dating has shown that these parts of the Cedar Mountain Formation were deposited about 124 million years ago.[9] The type specimen, CEU 184v.86, is currently housed at the College of Eastern Utah Prehistoric Museum, although Brigham Young University, the depository of Jensen's finds, currently houses the largest collection of Utahraptor fossils.

The type species (and only known species of Utahraptor), Utahraptor ostrommaysorum, was named by Kirkland, Gaston and Burge in 1993 for the American paleontologist John Ostrom, from Yale University's Peabody Museum of Natural History, and Chris Mays, of Dinamation International. Originally in the specific name the singular genitive ostrommaysi was used but in 2000 this was emended by George Olshevsky to the plural.[10] Earlier, it had been intended to name the species "Utahraptor spielbergi" after film director Steven Spielberg, in exchange for funding palaeontological research, but no agreement could be reached on the amount of financial assistance.[11]

A speculative reconstruction of Utahraptor lifestyles and behavior was presented in the 1999 BBC television series Walking with Dinosaurs.[12] The program portrayed Utahraptor with several anatomical inaccuracies including pronated hands and featherless skin, and depicted them living in Europe when the only fossils of Utahraptor have been found in western North America.[13]

References

  1. ^ a b c d e Kirkland, J.I.; Burge, D.; Gaston, R. (1993). "A large dromaeosaur [Theropoda] from the Lower Cretaceous of Utah". Hunteria. 2 (10): 1–16.
  2. ^ Kirkland, J.I. and Madsen, S.K. (2007). "The Lower Cretaceous Cedar Mountain Formation, eastern Utah: the view up an always interesting learning curve." Fieldtrip Guidebook, Geological Society of America, Rocky Mountain Section. 1-108 p.
  3. ^ Holtz, Thomas R. Jr. (2008) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages Supplementary Information
  4. ^ Turner, Alan H. (2007). "A basal dromaeosaurid and size evolution preceding avian flight". Science. 317 (5843): 1378–1381. doi:10.1126/science.1144066. PMID 17823350. {{cite journal}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
  5. ^ Xu, X.; Zhou, Z.; Wang, X.; Kuang, X.; Zhang, F.; Du, X. (2003). "Four-winged dinosaurs from China". Nature. 421 (6921): 335–340. doi:10.1038/nature01342. PMID 12540892.
  6. ^ Prum, R.; Brush, A.H. (2002). "The evolutionary origin and diversification of feathers". The Quarterly Review of Biology. 77 (3): 261–295. doi:10.1086/341993. PMID 12365352.
  7. ^ Turner, AH; Makovicky, PJ; Norell, MA (2007). "Feather quill knobs in the dinosaur Velociraptor". Science. 317 (5845): 1721. doi:10.1126/science.1145076. PMID 17885130.
  8. ^ Rothschild, B., Tanke, D. H., and Ford, T. L., 2001, Theropod stress fractures and tendon avulsions as a clue to activity: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, p. 331-336.
  9. ^ McDonald AT; Kirkland JI; DeBlieux DD; Madsen SK; Cavin J; et al. (2010). Farke, Andrew Allen (ed.). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLoS ONE. 5 (11): e14075. doi:10.1371/journal.pone.0014075. PMC 2989904. PMID 21124919. {{cite journal}}: Unknown parameter |author-separator= ignored (help)CS1 maint: unflagged free DOI (link)
  10. ^ Olshevsky, G., 2000, An annotated checklist of dinosaur species by continent. Mesozoic Meanderings 3: 1-157
  11. ^ Brooke Adams, 1993, "Director Loses Utahraptor Name Game", Deseret News, 15 June 1993
  12. ^ "Unnatural history? Deconstructing the Walking with Dinosaurs phenomenon." Media Culture & Society, 25(3): 315-332. doi:10.1177/0163443703025003002
  13. ^ Haines, T. (2000). Walking with Dinosaurs: A Natural History. London: BBC Publishing.