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Plotopteridae

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Plotopterids
Temporal range: EoceneMiocene
Copepteryx
Scientific classification
Kingdom:
Phylum:
Class:
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Plotopteridae

Howard, 1969
Genera

Copepteryx
Hokkaidornis
Phocavis
Plotopterum
Tonsala

Plotopteridae is the name of an extinct family of flightless seabirds from the order Pelecaniformes. Related to the gannets and boobies, they exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant penguins. That they lived in the North Pacific, the other side of the world from the penguins, has led to them being described at times as the Northern Hemisphere's penguins, though they were not closely related. More recent studies have shown, however, that the shoulder-girdle, forelimb and sternum of plotopterids differ significantly from those of penguins, so comparisons in terms of function may not be entirely accurate.[1]

Their fossils have been found in California, Washington and Japan. They ranged in size from that of a large cormorant (such as a Brandt's cormorant), to being 2 m long. They had shortened wings designed for underwater wing-propelled pursuit diving (like penguins or the now extinct great auk), a body skeleton similar to that of the darter and the skull similar to that of a sulid.

Tonsala hildegardae fossils

The earliest known Plotopteridae species, Phocavis maritimus lived in the mid-Eocene, but most of the known species lived in the early and mid-Miocene, after which it appears they went extinct. That they went extinct at the same time as the giant penguins of the Southern Hemisphere, which also coincided with the radiation of the seals and dolphins, has led to speculation that the expansion of marine mammals was responsible for the extinction of the Plotopteridae, though this has not been formally tested.

References

  • Howard, H. (1969). "A new avian fossil from Kern County, California" (PDF). Condor. 71: 68–69. doi:10.2307/1366050.
  • Olson, Storrs L.; Hasegawa, Yoshikazu; Hasegawa (1979). "Fossil Counterparts of Giant Penguins from the North Pacific". Science. 206 (4419): 688–689. Bibcode:1979Sci...206..688O. doi:10.1126/science.206.4419.688.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  • Olson, Storrs L. & Hasegawa, Yoshikazu (1996). "A new genus and two new species of gigantic Plotopteridae from Japan (Aves: Pelecaniformes)". J. Vert. Paleontol. 16 (4): 742–751. doi:10.1080/02724634.1996.10011362.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  • Schreiber, E.A. & Burger, J. (2001). Biology of Marine Birds. ISBN 0-8493-9882-7.{{cite book}}: CS1 maint: multiple names: authors list (link)
  • Mayr, Gerald (2005). "Tertiary plotopterids (Aves, Plotopteridae) and a novel hypothesis on the phylogenetic relationships of penguins (Spheniscidae)" (PDF). Journal of Zoological Systematics. 43 (1): 67–71. doi:10.1111/j.1439-0469.2004.00291.x.
  1. ^ Tatsuro et al, New Skeletal Remains of Plotopterids from Japan, SVP 2015