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{{Use dmy dates|date=October 2012}}
{{Redirect|Animalia}}
{{Other uses}}
{{pp-semi-protected|small=yes}}
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{{Taxobox
| color = {{taxobox color|[[animalia]]}}
| name = Animals
| fossil_range = [[Ediacaran]] – Recent {{fossilrange|610|0|}}
| image = Animal diversity.png
| image_width = 250px
| domain = [[Eukaryota]]
{{Taxobox_norank_entry | taxon = [[Opisthokonta]]}}
{{Taxobox_norank_entry | taxon = [[Holozoa]]}}
{{Taxobox_norank_entry | taxon = [[Filozoa]]}}
| regnum = '''Animalia'''
| regnum_authority = [[Carolus Linnaeus|Linnaeus]], [[Systema Naturae|1758]]
| subdivision_ranks = [[Phylum|Phyla]]
| subdivision =
* '''Subkingdom [[Parazoa]]'''
** [[Sponge|Porifera]]
** [[Placozoa]]
* '''Subkingdom [[Eumetazoa]]'''
** '''[[Radiata]] (unranked)'''
*** [[Ctenophora]]
*** [[Cnidaria]]
** '''[[Bilateria]] (unranked)'''
*** [[Orthonectida]]
*** [[Rhombozoa]]
*** [[Acoelomorpha]]
*** [[Chaetognatha]]
*** '''Superphylum [[Deuterostomia]]'''
**** [[Chordata]]
**** [[Hemichordata]]
**** [[Echinoderm]]ata
**** [[Xenoturbellida]]
**** [[Vetulicolia]] [[extinction|†]]
*** '''[[Protostomia]] (unranked)'''
**** '''Superphylum [[Ecdysozoa]]'''
***** [[Kinorhyncha]]
***** [[Loricifera]]
***** [[Priapulida]]
***** [[Nematoda]]
***** [[Nematomorpha]]
***** [[Lobopodia]]
***** [[Onychophora]]
***** [[Tardigrada]]
***** [[Arthropoda]]
**** '''Superphylum [[Platyzoa]]'''
***** [[Platyhelminthes]]
***** [[Gastrotricha]]
***** [[Rotifera]]
***** [[Acanthocephala]]
***** [[Gnathostomulida]]
***** [[Micrognathozoa]]
***** [[Cycliophora]]
**** '''Superphylum [[Lophotrochozoa]]'''
***** [[Sipuncula]]
***** [[Hyolitha]] [[extinction|†]]
***** [[Nemertea]]
***** [[Phoronida]]
***** [[Bryozoa]]
***** [[Entoprocta]]
***** [[Brachiopoda]]
***** [[Mollusca]]
***** [[Annelida]]
***** [[Echiura]]
}}

'''Animals''' are a major group of [[multicellular]], [[eukaryotic]] [[organism]]s of the [[Kingdom (biology)|kingdom]] '''Animalia''' or '''Metazoa'''. Their [[body plan]] eventually becomes fixed as they [[Developmental biology|develop]], although some undergo a process of [[metamorphosis]] later on in their life. Most animals are [[Motility|motile]], meaning they can move spontaneously and independently. All animals are also [[heterotroph]]s, meaning they must ingest other organisms or their products for [[sustenance]].

Most known animal [[phylum|phyla]] appeared in the fossil record as marine species during the [[Cambrian explosion]], about 542 million years ago.

==Etymology==
The word "animal" comes from the [[Latin]] word ''animalis'', meaning "having [[breath]]".<ref>{{cite book|last=Cresswell|first=Julia|title=The Oxford Dictionary of Word Origins|year=2010|publisher=Oxford University Press|location=New York|edition=2|isbn=978-0-19-954793-7|quote= ‘having the breath of life’, from anima ‘air, breath, life’ .}}</ref> In everyday colloquial usage, the word often refers to non-human members of kingdom Animalia. Sometimes, only closer relatives of humans such as [[mammal]]s and other [[vertebrate]]s are meant in colloquial use.<ref>{{cite web |url=http://m-w.com/dictionary/animals |title=Animals |publisher=[[Merriam-Webster's]] |quote=2 a : one of the lower animals as distinguished from human beings b : mammal; ''broadly'' : vertebrate |accessdate=16 May 2010}}</ref> The biological definition of the word refers to all members of the kingdom Animalia, encompassing creatures as diverse as sponges, jellyfish, insects and humans.<ref name="americanheritage_animal">{{cite encyclopedia |year=2006 |title=Animal |encyclopedia=The American Heritage Dictionary |publisher=Houghton Mifflin Company |edition=Forth }}</ref>

==Characteristics==
Animals have several characteristics that set them apart from other living things. Animals are [[eukaryote|eukaryotic]] and mostly [[multicellular]],<ref name="NationalZoo">{{cite web|url=http://nationalzoo.si.edu/Animals/GiantPandas/PandasForKids/classification/classification.htm|author=National Zoo|title=Panda Classroom|accessdate=30 September 2007| archiveurl= http://web.archive.org/web/20070929094936/http://nationalzoo.si.edu/Animals/GiantPandas/PandasForKids/classification/classification.htm| archivedate= 29 September 2007 <!--DASHBot-->| deadurl= no}}</ref> which separates them from [[bacteria]] and most [[protist]]s. They are [[heterotroph]]ic,<ref name="Windows">{{cite web|url=http://www.windows.ucar.edu/tour/link=/earth/Life/heterotrophs.html&edu=high|author=Jennifer Bergman|title=Heterotrophs|accessdate=30 September 2007| archiveurl= http://web.archive.org/web/20070829051950/http://www.windows.ucar.edu/tour/link=/earth/Life/heterotrophs.html&edu=high| archivedate= 29 August 2007 <!--DASHBot-->| deadurl= no}}</ref> generally digesting food in an internal chamber, which separates them from plants and [[alga]]e.<ref>{{cite journal |author=Douglas AE, Raven JA |title=Genomes at the interface between bacteria and organelles |journal=Philosophical transactions of the Royal Society of London. Series B, Biological sciences |volume=358 |issue=1429 |pages=5–17; discussion 517–8 |year=2003|pmid=12594915 |pmc=1693093 |doi=10.1098/rstb.2002.1188 |first1=AE |first2=JA}}</ref> They are also distinguished from plants, algae, and [[fungus|fungi]] by lacking rigid [[cell wall]]s.<ref name="AnimalCells">{{cite web|url=http://micro.magnet.fsu.edu/cells/animalcell.html|author=Davidson, Michael W.|title=Animal Cell Structure|accessdate=20 September 2007| archiveurl= http://web.archive.org/web/20070920235924/http://micro.magnet.fsu.edu/cells/animalcell.html| archivedate= 20 September 2007 <!--DASHBot-->| deadurl= no}}</ref> All animals are [[motility|motile]],<ref name="Concepts">{{cite web|url=http://employees.csbsju.edu/SSAUPE/biol116/Zoology/digestion.htm|author=Saupe, S.G|title=Concepts of Biology|accessdate=30 September 2007}}</ref> if only at certain life stages. In most animals, [[embryo]]s pass through a [[Blastula|blastula stage]],<ref>{{cite book|last=Minkoff|first=Eli C.|title=Barron's EZ-101 Study Keys Series: Biology|year=2008|publisher=Barron's Educational Series|isbn=978-0-7641-3920-8|edition=2, revised|page=48}}</ref> which is a characteristic exclusive to animals.

===Structure===
With a few exceptions, most notably the [[sponge]]s (Phylum [[Porifera]]) and [[Placozoa]], animals have [[body|bodies]] differentiated into separate [[biological tissue|tissues]]. These include [[muscle]]s, which are able to contract and control locomotion, and [[nervous system|nerve tissues]], which send and process signals. Typically, there is also an internal [[digestion|digestive]] chamber, with one or two openings.<ref name=scienceperspectives10/> Animals with this sort of organization are called metazoans, or [[eumetazoan]]s when the former is used for animals in general.<ref>{{cite book|title=Fossil Invertebrates|year=1983|publisher=CUP Archive|isbn=978-0-521-27028-1|page=350|author=Gero HIllmer|coauthors=Ulrich Lehmann|page=54}}</ref>

All animals have [[eukaryotic]] cells, surrounded by a characteristic [[extracellular matrix]] composed of [[collagen]] and elastic [[glycoprotein]]s.<ref>{{cite book|title=Molecular Biology of the Cell|year=2002|publisher=Garland Science|location=New York|url=http://www.ncbi.nlm.nih.gov/books/NBK26810/|author=Alberts, Bruce|edition=4|coauthors=Alexander Johnson, Julian Lewis, Martin Raff, Keith Roberts, and Peter Walter}}</ref> This may be calcified to form structures like [[animal shell|shells]], [[bone]]s, and [[spicule]]s.<ref>{{cite book|last=Sangwal|first=Keshra|title=Additives and crystallization processes: from fundamentals to applications|year=2007|publisher=John Wiley and Sons|isbn=978-0-470-06153-4|page=212}}</ref> During development, it forms a relatively flexible framework<ref>{{cite book|last=Becker|first=Wayne M.|title=The world of the cell|year=1991|publisher=Benjamin/Cummings|isbn=978-0-8053-0870-9}}</ref> upon which cells can move about and be reorganized, making complex structures possible. In contrast, other [[multicellular organism]]s, like plants and fungi, have cells held in place by cell walls, and so develop by progressive growth.<ref name=scienceperspectives10/> Also, unique to animal cells are the following intercellular junctions: [[tight junction]]s, [[gap junction]]s, and [[desmosome]]s.<ref>{{cite book|last=Magloire|first=Kim|title=Cracking the AP Biology Exam, 2004–2005 Edition|year=2004|publisher=The Princeton Review|isbn=978-0-375-76393-9|page=45}}</ref>

===Reproduction and development===
{{see also|Sexual reproduction#Animals|Asexual reproduction#Examples in animals}}
[[File:Mitosis-fluorescent.jpg|thumb|left|A [[newt]] [[lung]] [[cell (biology)|cell]] [[staining (biology)|stained]] with [[fluorescent]] [[dye]]s undergoing the early [[anaphase]] stage of [[mitosis]]]]

Nearly all animals undergo some form of [[Sexual reproduction#Animals|sexual reproduction]].<ref>{{cite book|last=Knobil|first=Ernst|title=Encyclopedia of reproduction, Volume 1|year=1998|publisher=Academic Press|isbn=978-0-12-227020-8|page=315}}</ref> They have a few specialized [[Gamete|reproductive cells]], which undergo [[meiosis]] to produce smaller, motile [[spermatozoon|spermatozoa]] or larger, non-motile [[ovum|ova]].<ref>{{cite book|last=Schwartz|first=Jill|title=Master the GED 2011 (w/CD)|year=2010|publisher=Peterson's|isbn=978-0-7689-2885-3|page=371}}</ref> These fuse to form [[zygote]]s, which develop into new individuals.<ref>{{cite book|last=Hamilton|first=Matthew B.|title=Population genetics|year=2009|publisher=Wiley-Blackwell|isbn=978-1-4051-3277-0|page=55}}</ref>

Many animals are also capable of [[asexual reproduction]].<ref>{{cite book|title=Reproductive Biology of Invertebrates, Progress in Asexual Reproduction, Volume 11|year=2002|publisher=Wiley|author=Adiyodi, K. G.|coauthors=Roger N. Hughes, Rita G. Adiyodi|page=116}}</ref> This may take place through [[parthenogenesis]], where fertile eggs are produced without mating, budding, or [[fragmentation (reproduction)|fragmentation]].<ref>{{cite book|last=Kaplan|title=GRE exam subject test|year=2008|publisher=Kaplan Publishing|isbn=978-1-4195-5218-2|page=233}}</ref>

A [[zygote]] initially develops into a hollow sphere, called a [[blastula]],<ref>{{cite book|last=Tmh|title=Study Package For Medical College Entrance Examinations|year=2006|publisher=Tata McGraw-Hill|isbn=978-0-07-061637-0|page=6.22}}</ref> which undergoes rearrangement and differentiation. In sponges, blastula larvae swim to a new location and develop into a new sponge.<ref>{{cite book|title=General zoology|year=1984|publisher=Saunders College Pub|isbn=978-0-03-062451-3|author=Ville, Claude Alvin|coauthors=Warren Franklin Walker, Robert D. Barnes|page=467}}</ref> In most other groups, the blastula undergoes more complicated rearrangement.<ref>{{cite book|title=Human embryology: (prenatal development of form and function)|year=1945|publisher=Williams & Wilkins|author=Hamilton, William James|coauthors=James Dixon Boyd, Harland Winfield Mossman|page=330}}</ref> It first [[invagination|invaginates]] to form a [[gastrula]] with a digestive chamber, and two separate [[germ layer]]s&nbsp;— an external [[ectoderm]] and an internal [[endoderm]].<ref>{{cite book|last=Philips|first=Joy B.|title=Development of vertebrate anatomy|year=1975|publisher=Mosby|isbn=978-0-8016-3927-2|page=176}}</ref> In most cases, a [[mesoderm]] also develops between them.<ref>{{cite book|title=The Encyclopedia Americana: a library of universal knowledge, Volume 10|year=1918|publisher=Encyclopedia Americana Corp.|page=281}}</ref> These germ layers then differentiate to form tissues and organs.<ref>{{cite book|title=The science of entomology|year=1998|publisher=WCB McGraw-Hill|isbn=978-0-697-22848-2|author=Romoser, William S.|coauthors=J. G. Stoffolano|page=156}}</ref>

===Food and energy sourcing===
{{Main|Animal nutrition}}

All animals are [[heterotroph]]s, meaning that they feed directly or indirectly on other living things.<ref>{{cite book|last=Rastogi|first=V. B.|title=Modern Biology|year=1997|publisher=Pitambar Publishing|isbn=978-81-209-0496-5|page=3}}</ref> They are often further subdivided into groups such as [[carnivore]]s, [[herbivore]]s, [[omnivore]]s, and [[parasite]]s.<ref>{{cite book|last=Levy|first=Charles K.|title=Elements of Biology|year=1973|publisher=Appleton-Century-Crofts|isbn=978-0-390-55627-1|page=108}}</ref>

[[Predation]] is a [[biological interaction]] where a predator (a heterotroph that is hunting) feeds on its prey (the organism that is attacked).<ref name="Ecology">Begon, M., Townsend, C., Harper, J. (1996). ''[[Ecology: Individuals, populations and communities]]'' (Third edition). Blackwell Science, London. ISBN 0-86542-845-X, ISBN 0-632-03801-2, ISBN 0-632-04393-8.</ref> Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of the prey.<ref>[http://www.britannica.com/search?query=predation predation]. Britannica.com. Retrieved on 2011-11-23.</ref> The other main category of consumption is [[detritivory]], the consumption of dead [[organic matter]].<ref>{{cite book|title=Geomorphology and environmental impact assessment|year=2001|publisher=Taylor & Francis|isbn=978-90-5809-344-8|author=Marchetti, Mauro|coauthors=Victoria Rivas|page=84}}</ref> It can at times be difficult to separate the two [[feeding behaviour]]s, for example, where [[parasite|parasitic species]] prey on a host organism and then lay their eggs on it for their offspring to feed on its decaying corpse. Selective pressures imposed on one another has led to an [[evolutionary arms race]] between prey and predator, resulting in various [[antipredator adaptations]].<ref>{{cite book|title=Ecology of marine fishes: California and adjacent waters|year=2006|publisher=University of California Press|isbn=978-0-520-24653-9|author=Allen, Larry Glen|coauthors=Daniel J. Pondella, Michael H. Horn|page=428}}</ref>

Most animals indirectly use the energy of [[sunlight]] by eating plants or plant-eating animals. Most plants use light to convert [[inorganic]] molecules in their environment into [[carbohydrate]]s, [[:fat]]s, [[protein]]s and other biomolecules, characteristically containing [[Reduction (chemistry)|reduced]] carbon in the form of carbon-hydrogen bonds. Starting with [[carbon dioxide]] (CO<sub>2</sub>) and water (H<sub>2</sub>O), photosynthesis converts the energy of sunlight into chemical energy in the form of simple sugars (e.g., [[glucose]]), with the release of molecular [[oxygen]]. These sugars are then used as the building blocks for plant growth, including the production of other biomolecules.<ref name=scienceperspectives10>{{cite book|title=Science Perspectives 10|year=2010|publisher=Nelson Education Ltd.|isbn=978-0-17-635528-9|last1=Adam-Carr|first1=Christine|last2=Hayhoe|first2=Christy|last3=Hayhoe|first3=Douglas|last4=Hayhoe|first4=Katharine}}</ref> When an animal eats plants (or eats other animals which have eaten plants), the reduced carbon compounds in the food become a source of energy and building materials for the animal.<ref>{{cite book|title=Understanding Science: Upper Primary|last=Clutterbuck|first=Peter|page=9|year=2000|publisher=Blake Education|isbn=978-1-86509-170-9}}</ref> They are either used directly to help the animal grow, or broken down, releasing stored solar energy, and giving the animal the energy required for motion.<ref>{{cite book|last=Gupta|first=P.K.|title=Genetics Classical To Modern|publisher=Rastogi Publications|isbn=978-81-7133-896-2|page=26}}</ref><ref>{{cite book|last1=Garrett|first1=Reginald|last2=Grisham|first2=Charles M.|title=Biochemistry|year=2010|publisher=Cengage Learning|isbn=978-0-495-10935-8|page=535}}</ref>

Animals living close to [[hydrothermal vent]]s and [[cold seep]]s on the [[Seabed|ocean floor]] are not dependent on the energy of sunlight.<ref>{{cite journal|year=1996|journal=New scientist|publisher=IPC Magazines|volume=152|issue=2050–2055|page=105}}</ref> Instead [[chemosynthesis|chemosynthetic]] [[archaea]] and bacteria form the base of the [[food chain]].<ref>{{cite book|last=Castro|first=Peter|coauthors=Michael E. Huber|title=Marine Biology|publisher=McGraw-Hill|year=2007|edition=7|page=376|isbn=978-0-07-722124-9}}</ref>

==Origin and fossil record==
{{See|Urmetazoan}}
[[File:Dunkleosteus BW.jpg|thumb|''[[Dunkleosteus]]'' was a gigantic, {{convert|10|m|ft|adj=mid|-long}} [[prehistoric fish]].<ref>[http://www.smh.com.au/news/science/jaws-of-steel-on-this-fish-tank/2006/11/29/1164777657728.html Monster fish crushed opposition with strongest bite ever], smh.com.au</ref>]]
[[File:Vernanimalcula.jpg|thumb|''[[Vernanimalcula|Vernanimalcula guizhouena]]'' is a fossil believed by some to represent the earliest known member of the [[Bilateria]].]]
Animals are generally considered to have [[Evolution|evolved]] from a [[flagellate]]d eukaryote.<ref>{{cite book|last=Campbell|first=Niel A.|title=Biology|year=1990|publisher=Benjamin/Cummings Pub. Co.|isbn=978-0-8053-1800-5|edition=2|page=560}}</ref> Their closest known living relatives are the [[choanoflagellate]]s, collared flagellates that have a morphology similar to the choanocytes of certain sponges.<ref>{{cite book|title=Emerging model organisms: a laboratory manual, Volume 1|year=2008|publisher=Cold Spring Harbor Laboratory Press|isbn=978-0-87969-872-0|page=592|edition=illustrated|editor=Richard R. Behringer, Alexander D. Johnson, Robert E. Krumlauf, Michael K. Levine, Nipam Patel, Neelima Sinha|page=1}}</ref> [[Molecular]] studies place animals in a supergroup called the [[opisthokont]]s, which also include the choanoflagellates, [[fungus|fungi]] and a few small parasitic [[protist]]s.<ref>{{cite book|title=Strickberger's evolution: the integration of genes, organisms and populations|year=2008|publisher=Jones & Bartlett Learning|isbn=978-0-7637-0066-9|author=Hall, Brian Keith|coauthors=Benedikt Hallgrímsson, Monroe W. Strickberger|page=278}}</ref> The name comes from the posterior location of the [[flagellum]] in motile cells, such as most animal spermatozoa, whereas other [[eukaryotes]] tend to have anterior flagella.<ref>{{cite book|last=Hamilton|first=Gina|title=Kingdoms of Life – Animals (ENHANCED eBook)|publisher=Lorenz Educational Press|isbn=978-1-4291-1610-7|page=9}}</ref>

The first fossils that might represent animals appear in the Trezona Formation at Trezona Bore, West Central Flinders, South Australia.<ref name="roseMaloof">{{cite journal | author1= Maloof, Adam C. |author2= Rose, Catherine V. |author3= Beach, Robert |author4= Samuels, Bradley M. |author5= Calmet, Claire C. |author6= Erwin, Douglas H. |author7= Poirier, Gerald R. |author8= Yao, Nan |author9= Simons, Frederik J.|title= Possible animal-body fossils in pre-Marinoan limestones from South Australia |journal= Nature Geoscience | volume= 3 | page= 653 |date= 17 August 2010 |url= http://www.nature.com/ngeo/journal/vaop/ncurrent/full/ngeo934.html |doi=10.1038/ngeo934 | issue= 9|bibcode = 2010NatGe...3..653M }}</ref> These fossils are interpreted as being early sponges. They were found in 665-million-year-old rock.<ref name="roseMaloof"/>

The next oldest possible animal fossils are found towards the end of the [[Precambrian]], around 610 million years ago, and are known as the [[Ediacaran biota|Ediacaran or Vendian biota]].<ref>{{cite book|title=The annotated Origin: a facsimile of the first edition of On the origin of species|year=2009|publisher=Harvard University Press|isbn=978-0-674-03281-1|author=Costa, James T.|coauthors=Charles Darwin|page=308}}</ref> These are difficult to relate to later fossils, however. Some may represent precursors of modern phyla, but they may be separate groups, and it is possible they are not really animals at all.<ref>{{cite book|last=Schopf|first=J. William|title=Evolution!: facts and fallacies|year=1999|publisher=Academic Press|isbn=978-0-12-628860-5|page=7}}</ref>

Aside from them, most known animal phyla make a more or less simultaneous appearance during the [[Cambrian]] period, about 542 million years ago.<ref>{{cite book|title=Fossils at a Glance|year=2009|publisher=John Wiley and Sons|isbn=978-1-4051-9336-8|author=Milsom, Clare|coauthors=Sue Rigby}}</ref> It is still disputed whether this event, called the [[Cambrian explosion]], represents a rapid divergence between different groups or a change in conditions that made fossilization possible.

Some paleontologists suggest that animals appeared much earlier than the Cambrian explosion, possibly as early as 1 billion years ago.<ref>{{cite book|title=Biology|year=2005|publisher=Pearson, Benjamin Cummings|isbn=978-0-8053-7171-0|author=Campbell. Neil A.|edition=7|coauthors=Jane B. Reece|page=526}}</ref> [[Trace fossil]]s such as tracks and burrows found in the [[Tonian]] era indicate the presence of [[triploblastic]] worms, like metazoans, roughly as large (about 5&nbsp;mm wide) and complex as [[earthworms]].<ref name="Seilacher1998">{{cite journal
| title=Animals More Than 1 Billion Years Ago: Trace Fossil Evidence from India
| journal=Science
| volume=282
| pages=80–83
| year=1998
| author=Seilacher, A., Bose, P.K. and Pflüger, F.
| doi=10.1126/science.282.5386.80
| pmid=9756480
| first1=A
| first2=PK
| first3=F
| issue=5386
|bibcode = 1998Sci...282...80S }}</ref> During the beginning of the Tonian period around 1 billion years ago, there was a decrease in [[Stromatolite]] diversity, which may indicate the appearance of grazing animals, since stromatolite diversity increased when grazing animals went extinct at the [[Permian–Triassic extinction event|End Permian]] and [[Ordovician–Silurian extinction event|End Ordovician]] extinction events, and decreased shortly after the grazer populations recovered. However the discovery that tracks very similar to these early trace fossils are produced today by the giant single-celled protist ''[[Gromia sphaerica]]'' casts doubt on their interpretation as evidence of early animal evolution.<ref name="Matz2008">{{cite journal | last = Matz | first = Mikhail V. | coauthors = Tamara M. Frank, N. Justin Marshall, Edith A. Widder and Sonke Johnsen | title = Giant Deep-Sea Protist Produces Bilaterian-like Traces | journal = Current Biology | volume = 18| issue = 23 | pages = 1–6 |year = 2008| url = http://www.biology.duke.edu/johnsenlab/pdfs/pubs/sea%20grapes%202008.pdf | doi = 10.1016/j.cub.2008.10.028 | accessdate = 2008-12-05 | pmid = 19026540 | last1 = Matz | first1 = MV | last2 = Frank | first2 = TM | last3 = Marshall | first3 = NJ | last4 = Widder | first4 = EA | last5 = Johnsen | first5 = S | archiveurl= http://web.archive.org/web/20081216211211/http://www.biology.duke.edu/johnsenlab/pdfs/pubs/sea%20grapes%202008.pdf| archivedate= 16 December 2008 <!--DASHBot-->| deadurl= no}}</ref><ref name="MSNBC200811">{{cite news | last = Reilly | first = Michael | title = Single-celled giant upends early evolution | publisher = MSNBC | date = 20 November 2008 | url = http://www.msnbc.msn.com/id/27827279/ | accessdate = 2008-12-05}}</ref>

It has been estimated that 99.9% of animals that have ever existed are [[extinct]].<ref>[http://www.bbc.co.uk/lastchancetosee/sites/about/extinction.shtml The thing about extinction] retrieved 20 October 2012</ref>

==Groups of animals==
[[File:AnimalsRelativeNumbers.png|thumb|301px|right|The relative number of species contributed to the total by each phylum of animals.]]

===Porifera, Radiata and basal Bilateria===

Phylogenetic analysis suggests that the [[Porifera]] and [[Ctenophora]] diverged before a clade that gave rise to the [[Bilateria]], [[Cnidaria]] and [[Placozoa]].<ref name=Ryan2010>Ryan JF, Pang K; NISC Comparative Sequencing Program, Mullikin JC, Martindale MQ, Baxevanis AD (2010) The homeodomain complement of the ctenophore ''Mnemiopsis leidyi'' suggests that Ctenophora and Porifera diverged prior to the ParaHoxozoa. ''Evodevo'' 1(1): 9</ref> Another study based on the presence/absence of [[intron]]s suggests that Cnidaria, Porifera and Placozoa may be a sister group of Bilateria and Ctenophora.<ref name=Lehmann2012>Lehmann J, Stadler PF, Krauss V (2012) Near intron pairs and the metazoan tree. Mol Phylogenet Evol pii: S1055-7903(12)00444-7. doi: 10.1016/j.ympev.2012.11.012 </ref>

[[File:Elephant-ear-sponge.jpg|thumb|left|Orange elephant ear sponge, ''[[Agelas clathrodes]]'', in foreground. Two corals in the background: a [[sea fan]], ''[[Iciligorgia schrammi]]'', and a sea rod, ''[[Plexaurella nutans]]''.]]
The sponges ([[Porifera]]) were long thought to have diverged from other animals early.<ref>{{cite book|title=An Introduction to Porifera|year=2003|publisher=Anmol Publications PVT. LTD.|isbn=978-81-261-0675-2|author=Bhamrah, H. S.|coauthors=Kavita Juneja|page=58}}</ref> They lack the complex organization found in most other phyla.<ref>{{cite book|last=Sumich|first=James L.|title=Laboratory and Field Investigations in Marine Life|year=2008|publisher=Jones & Bartlett Learning|isbn=978-0-7637-5730-4|page=67}}</ref> Their cells are differentiated, but in most cases not organized into distinct tissues.<ref>{{cite book|last=Jessop|first=Nancy Meyer|title=Biosphere; a study of life|year=1970|publisher=Prentice-Hall|page=428}}</ref> Sponges typically feed by drawing in water through pores.<ref>{{cite book|last=Sharma|first=N. S.|title=Continuity And Evolution Of Animals|year=2005|publisher=Mittal Publications|isbn=978-81-8293-018-6|page=106}}</ref> [[Archaeocyatha]], which have fused skeletons, may represent sponges or a separate phylum.<ref>{{cite book|title=McGraw-Hill encyclopedia of science & technology: MET-NIC., Volume 11|year=1997|publisher=McGraw-Hill|isbn=978-0-07-911504-1|page=809|url=http://books.google.com/?id=GS5YAAAAMAAJ|edition=8|accessdate=19 March 2011|page=59}}</ref> However, a phylogenomic study in 2008 of 150 genes in 29 animals across 21 phyla revealed that it is the [[Ctenophora]] or comb jellies which are the basal lineage of animals, at least among those 21 phyla. The authors speculate that sponges—or at least those lines of sponges they investigated—are not so primitive, but may instead be secondarily simplified.<ref name=class>{{cite journal|title=Broad phylogenomic sampling improves resolution of the animal tree of life|doi=10.1038/nature06614|pmid=18322464 | volume=452|issue=7188|year=2008|month=April|journal=Nature|pages=745–9|author=Dunn CW, Hejnol A, Matus DQ, ''et al.''|bibcode = 2008Natur.452..745D }}</ref>

Among the other phyla, the Ctenophora and the [[Cnidaria]], which includes [[sea anemone]]s, [[coral]]s, and [[jellyfish]], are radially symmetric and have digestive chambers with a single opening, which serves as both the mouth and the anus.<ref>{{cite book|title=A living bay: the underwater world of Monterey Bay|year=2000|publisher=University of California Press|isbn=978-0-520-22149-9|author=Langstroth, Lovell|coauthors=Libby Langstroth, Todd Newberry, Monterey Bay Aquarium|page=244}}</ref> Both have distinct tissues, but they are not organized into [[organ (anatomy)|organs]].<ref>{{cite book|last=Safra|first=Jacob E.|title=The New Encyclopædia Britannica, Volume 16|year=2003|publisher=Encyclopædia Britannica|isbn=978-0-85229-961-6|page=523}}</ref> There are only two main germ layers, the ectoderm and endoderm, with only scattered cells between them. As such, these animals are sometimes called [[diploblastic]].<ref>{{cite book|last=Kotpal|first=R. L.|title=Modern Text Book of Zoology: Invertebrates|publisher=Rastogi Publications|isbn=978-81-7133-903-7|page=184}}</ref> The tiny [[placozoan]]s are similar, but they do not have a permanent digestive chamber.

The remaining animals form a [[monophyletic]] group called the [[Bilateria]]. For the most part, they are [[Symmetry (biology)|bilaterally symmetric]], and often have a specialized head with feeding and sensory organs. The body is [[triploblastic]], i.e. all three germ layers are well-developed, and tissues form distinct organs. The digestive chamber has two openings, a mouth and an anus, and there is also an internal body cavity called a [[coelom]] or pseudocoelom. There are exceptions to each of these characteristics, however&nbsp;— for instance adult [[echinoderm]]s are radially symmetric, and certain [[parasitic worm]]s have extremely simplified body structures.

Genetic studies have considerably changed our understanding of the relationships within the Bilateria. Most appear to belong to two major lineages: the [[deuterostome]]s and the [[protostome]]s, the latter of which includes the [[Ecdysozoa]], [[Platyzoa]], and [[Lophotrochozoa]]. In addition, there are a few small groups of bilaterians with relatively similar structure that appear to have diverged before these major groups. These include the [[Acoelomorpha]], [[Rhombozoa]], and [[Orthonectida]]. The [[Myxozoa]], single-celled parasites that were originally considered Protozoa, are now believed to have developed from the Medusozoa as well.

===Deuterostomes===
[[File:Superbfairywrenscropped.jpeg|thumb|[[Superb Fairy-wren]], ''Malurus cyaneus'']]
[[Deuterostome]]s differ from the other Bilateria, called [[protostome]]s, in several ways. In both cases there is a complete digestive tract. However, in protostomes, the first opening of the gut to appear in embryological development (the [[archenteron]]) develops into the mouth, with the anus forming secondarily. In deuterostomes the anus forms first, with the mouth developing secondarily.<ref>{{cite book|title=The Biomarker Guide: Biomarkers and isotopes in petroleum systems and Earth history|year=2005|publisher=Cambridge University Press|isbn=978-0-521-83762-0|author=Peters, Kenneth E.|coauthors=Clifford C. Walters, J. Michael Moldowan|page=717}}</ref> In most protostomes, cells simply fill in the interior of the gastrula to form the mesoderm, called schizocoelous development, but in deuterostomes, it forms through [[invagination]] of the endoderm, called enterocoelic pouching.<ref>{{cite book|last=Safra|first=Jacob E.|title=The New Encyclopædia Britannica, Volume 1; Volume 3|year=2003|publisher=Encyclopædia Britannica|isbn=978-0-85229-961-6|page=767}}</ref> Deuterostome embryos undergo radial [[Cleavage (embryo)|cleavage]] during cell division, while protostomes undergo spiral cleavage.<ref>{{cite journal|last=Valentine|first=James W.|date=July 1997|title=Cleavage patterns and the topology of the metazoan tree of life|journal=PNAS|publisher=The National Academy of Sciences|volume=94|pages=8001–8005|bibcode = 1997PNAS...94.8001V |doi = 10.1073/pnas.94.15.8001|pmid=9223303|pmc=21545}}</ref>

All this suggests the deuterostomes and protostomes are separate, monophyletic lineages. The main phyla of deuterostomes are the [[Echinodermata]] and [[Chordate|Chordata]].<ref>{{cite book|last=Hyde|first=Kenneth|title=Zoology: An Inside View of Animals|year=2004|publisher=Kendall Hunt|isbn=978-0-7575-0997-1|page=345}}</ref> The former are radially symmetric and exclusively marine, such as [[starfish]], [[sea urchin]]s, and [[Holothuroidea|sea cucumbers]].<ref>{{cite book|last=Alcamo|first=Edward|title=Biology Coloring Workbook|year=1998|publisher=The Princeton Review|isbn=978-0-679-77884-4|page=220}}</ref> The latter are dominated by the [[vertebrate]]s, animals with backbones.<ref>{{cite book|last=Holmes|first=Thom|title=The First Vertebrates|page=64|publisher=Infobase Publishing|year=2008|isbn=978-0-8160-5958-4}}</ref> These include [[:fish]], [[amphibian]]s, [[reptile]]s, [[:bird]]s, and [[mammal]]s.<ref>{{cite book|last=Rice|first=Stanley A.|title=Encyclopedia of evolution|publisher=Infobase Publishing|year=2007|page=75|isbn=978-0-8160-5515-9}}</ref>

In addition to these, the deuterostomes also include the [[Hemichordata]], or acorn worms.<ref>{{cite book|title=Asking about life|year=2005|publisher=Cengage Learning|isbn=978-0-534-40653-0|author=Tobin, Allan J.|coauthors=Jennie Dusheck|page=497}}</ref> Although they are not especially prominent today, the important fossil [[graptolite]]s may belong to this group.<ref>{{cite book|last=Safra|first=Jacob E.|title=The New Encyclopædia Britannica, Volume 19|year=2003|publisher=Encyclopædia Britannica|isbn=978-0-85229-961-6|page=791}}</ref>

The [[Chaetognatha]] or arrow worms may also be deuterostomes, but more recent studies suggest protostome affinities.

===Ecdysozoa===
[[File:Sympetrum flaveolum - side (aka).jpg|thumb|[[Yellow-winged darter]], ''Sympetrum flaveolum'']]
The [[Ecdysozoa]] are protostomes, named after the common trait of growth by moulting or [[ecdysis]].<ref>{{cite book|last=Dawkins|first=Richard|title=The Ancestor's Tale: A Pilgrimage to the Dawn of Evolution|year=2005|publisher=Houghton Mifflin Harcourt|isbn=978-0-618-61916-0|page=381}}</ref> The largest animal phylum belongs here, the [[Arthropoda]], including insects, spiders, crabs, and their kin. All these organisms have a body divided into repeating segments, typically with paired appendages. Two smaller phyla, the [[Onychophora]] and [[Tardigrada]], are close relatives of the arthropods and share these traits.

The ecdysozoans also include the [[Nematoda]] or roundworms, perhaps the second largest animal phylum. Roundworms are typically microscopic, and occur in nearly every environment where there is water.<ref>{{cite book|title=BioInquiry: making connections in biology|year=2003|publisher=John Wiley|isbn=978-0-471-20228-8|author=Prewitt, Nancy L.|coauthors=Larry S. Underwood, William Surver|page=289}}</ref> A number are important parasites.<ref>{{cite book|title=Parasites in social insects|year=1998|publisher=Princeton University Press|isbn=978-0-691-05924-2|last=Schmid-Hempel|first=Paul|page=75}}</ref> Smaller phyla related to them are the [[Nematomorpha]] or horsehair worms, and the [[Kinorhyncha]], [[Priapulida]], and [[Loricifera]]. These groups have a reduced coelom, called a pseudocoelom.

The remaining two groups of protostomes are sometimes grouped together as the [[Spiralia]], since in both embryos develop with [[spiral cleavage]].

===Platyzoa===
[[File:Bedford's Flatworm.jpg|thumb|''[[Pseudobiceros bedfordi]]'', (Bedford's flatworm)]]
The [[Platyzoa]] include the phylum [[Platyhelminthes]], the flatworms.<ref name=Gilson>{{cite book|last=Gilson|first=Étienne|title=El espíritu de la filosofía medieval|year=2004|publisher=Ediciones Rialp|isbn=978-84-321-3492-0|page=384}}</ref> These were originally considered some of the most primitive Bilateria, but it now appears they developed from more complex ancestors.<ref>{{cite journal |coauthors=Ruiz-Trillo, Iñaki; Riutort, Marta; Littlewood, D. Timothy J.; Herniou, Elisabeth A.; Baguñà, Jaume |year= 1999 |title=Acoel Flatworms: Earliest Extant Bilaterian Metazoans, Not Members of Platyhelminthes |journal=Science |volume=283 |issue=5409 |pages=1919–1923 |doi=10.1126/science.283.5409.1919 |accessdate= 2007-12-19 |author=Ruiz-Trillo, I. |pmid=10082465 |first1=I |first2=M |first3=DT |first4=EA |first5=J|bibcode = 1999Sci...283.1919R }}</ref> A number of [[parasites]] are included in this group, such as the [[Trematoda|flukes]] and [[tapeworm]]s.<ref name=Gilson/> Flatworms are [[Body cavity#No coelom (Acoelomate)|acoelomates]], lacking a body cavity, as are their closest relatives, the microscopic [[Gastrotricha]].<ref name="umodena">{{cite web |url=http://www.gastrotricha.unimore.it/overview.htm |title=Gastrotricha: Overview |accessdate=2008-01-26 |last=Todaro |first=Antonio |work=Gastrotricha: World Portal |publisher=University of Modena & Reggio Emilia}}</ref>

The other platyzoan phyla are mostly microscopic and [[Body cavity#Pseudocoelomates|pseudocoelomate]]. The most prominent are the [[Rotifera]] or rotifers, which are common in aqueous environments. They also include the [[Acanthocephala]] or spiny-headed worms, the [[Gnathostomulida]], [[Micrognathozoa]], and possibly the [[Cycliophora]].<ref name="IntroCyclio">{{cite journal |last=Kristensen |first= Reinhardt Møbjerg |year=2002 |title=An Introduction to Loricifera, Cycliophora, and Micrognathozoa |journal=Integrative and Comparative Biology |volume=42 |issue=3 |pages=641–651 |doi =10.1093/icb/42.3.641 |pmid=21708760}}</ref> These groups share the presence of complex jaws, from which they are called the [[Gnathifera (phylum)|Gnathifera]].

===Lophotrochozoa===
[[File:Grapevinesnail 01.jpg|thumb|[[Roman snail]], ''Helix pomatia'']]
The [[Lophotrochozoa]] include two of the most successful animal phyla, the [[Mollusca]] and [[Annelida]].<ref>{{cite web |url=http://www.lophelia.org/lophelia/biodiv_6.htm |title=Biodiversity: Mollusca|accessdate=2007-11-19 |publisher=The Scottish Association for Marine Science |archiveurl = http://web.archive.org/web/20060708083128/http://www.lophelia.org/lophelia/biodiv_6.htm |archivedate = 8 July 2006}}</ref><ref>{{cite video | people = Russell, Bruce J. (Writer), Denning, David (Writer) | title = Branches on the Tree of Life: Annelids| medium = VHS | publisher = BioMEDIA ASSOCIATES |year=2000 }}</ref> The former, which is the second-largest animal phylum by number of described species, includes animals such as [[snail]]s, [[clam]]s, and [[squid]]s, and the latter comprises the segmented worms, such as [[earthworm]]s and [[leech]]es. These two groups have long been considered close relatives because of the common presence of [[trochophore]] larvae, but the annelids were considered closer to the arthropods because they are both segmented.<ref>{{cite journal| coauthors=Eernisse, Douglas J.; Albert, James S.; Anderson , Frank E. | title=Annelida and Arthropoda are not sister taxa: A phylogenetic analysis of spiralean metazoan morphology | journal=Systematic Biology | volume=41 | issue=3 | pages = 305–330 | date=1 September 1992 | doi=10.2307/2992569 | author=Eernisse, Douglas J.| first=D. J.| first2=J. S.| first3=F. E.| jstor=2992569 }}</ref> Now, this is generally considered [[convergent evolution]], owing to many morphological and genetic differences between the two phyla.<ref>{{cite journal| coauthors=Kim, Chang Bae; Moon, Seung Yeo; Gelder, Stuart R.; Kim, Won | title=Phylogenetic Relationships of Annelids, Molluscs, and Arthropods Evidenced from Molecules and Morphology | journal=Journal of Molecular Evolution | volume=43 | issue=3 | pages = 207–215 | publisher = Springer | location = New York| year = 1996| doi = 10.1007/PL00006079| author=Eernisse, Douglas J.| pmid=8703086 }}</ref>

The Lophotrochozoa also include the [[Nemertea]] or ribbon worms, the [[Sipuncula]], and several phyla that have a ring of ciliated tentacles around the mouth, called a [[lophophore]].<ref>{{Cite book | url = http://www.ucmp.berkeley.edu/glossary/gloss7/lophophore.html | title= The Lophophore| author =Collins, Allen G. | year = 1995 | publisher = University of California Museum of Paleontology}}</ref> These were traditionally grouped together as the lophophorates.<ref>{{cite journal| coauthors=Adoutte, André; Balavoine, Guillaume; Lartillot, Nicolas; Lespinet, Olivier; Prud'homme, Benjamin; de Rosa, Renaud | title=The new animal phylogeny: Reliability and implications | journal=Proceedings of the National Academy of Sciences | volume=97 | issue=9 | pages = 4453–4456 | year= 2000 | pmid=10781043 | doi=10.1073/pnas.97.9.4453 | author=Adoutte, A.| first1=A| first2=G| first3=N| first4=O| first5=B| first6=R| pmc=34321 |bibcode = 2000PNAS...97.4453A }}</ref> but it now appears that the lophophorate group may be [[paraphyletic]],<ref>{{cite web | first = Yale J. | last = Passamaneck | contribution = Woods Hole Oceanographic Institution | title = Molecular Phylogenetics of the Metazoan Clade Lophotrochozoa | year = 2003 | page = 124 | url = http://handle.dtic.mil/100.2/ADA417356 | format = PDF }}</ref> with some closer to the nemerteans and some to the molluscs and annelids.<ref>{{cite journal| coauthors=Sundberg, Per; Turbevilleb, J. M.; Lindha, Susanne | title=Phylogenetic relationships among higher nemertean (Nemertea) taxa inferred from 18S rDNA sequences | journal=Molecular Phylogenetics and Evolution | volume=20 | issue=3 | pages = 327–334 | year = 2001 | doi = 10.1006/mpev.2001.0982 | author=Adoutte, A.| pmid=11527461| last1=Sundberg| first1=P| last2=Turbeville| first2=JM| last3=Lindh| first3=S}}</ref><ref>{{cite journal| coauthors=Boore, Jeffrey L.; Staton, Joseph L | title=The mitochondrial genome of the Sipunculid Phascolopsis gouldii supports its association with Annelida rather than Mollusca | journal=Molecular Biology and Evolution | volume=19 | issue=2 | pages = 127–137 | year = 2002 | url = http://mbe.oxfordjournals.org/cgi/reprint/19/2/127.pdf | format=PDF | pmid=11801741 | accessdate = 2007-11-19| last1=Boore| first1=JL| first2=JL | archiveurl= http://web.archive.org/web/20071128200059/http://mbe.oxfordjournals.org/cgi/reprint/19/2/127.pdf| archivedate= 28 November 2007 <!--DASHBot-->| deadurl= no}}</ref> They include the [[Brachiopoda]] or lamp shells, which are prominent in the fossil record, the [[Entoprocta]], the [[Phoronida]], and possibly the [[Bryozoa]] or moss animals.<ref>{{cite journal | last = Nielsen | first = Claus | year = 2001| title = Bryozoa (Ectoprocta: ‘Moss’ Animals) | journal = Encyclopedia of Life Sciences | publisher = John Wiley & Sons, Ltd | doi = 10.1038/npg.els.0001613 | url = http://mrw.interscience.wiley.com/emrw/9780470015902/els/article/a0001613/current/abstract | accessdate = 2008-01-19 }}</ref>

==Model organisms==
{{Main|Model organism|Animal testing}}
Because of the great diversity found in animals, it is more economical for scientists to study a small number of chosen species so that connections can be drawn from their work and conclusions extrapolated about how animals function in general. Because they are easy to keep and breed, the fruit fly ''[[Drosophila melanogaster]]'' and the nematode ''[[Caenorhabditis elegans]]'' have long been the most intensively studied metazoan [[model organism]]s, and were among the first life-forms to be genetically sequenced. This was facilitated by the severely reduced state of their [[genome]]s, but as many [[gene]]s, [[intron]]s, and [[genetic linkage|linkages]] lost, these ecdysozoans can teach us little about the origins of animals in general. The extent of this type of evolution within the superphylum will be revealed by the crustacean, annelid, and molluscan [[genome project]]s currently in progress. Analysis of the [[starlet sea anemone]] genome has emphasised the importance of sponges, placozoans, and [[choanoflagellate]]s, also being sequenced, in explaining the arrival of 1500 ancestral genes unique to the Eumetazoa.<ref>
{{cite journal
|author=N.H. Putnam, ''et al.''
|year=2007
|title=Sea anemone genome reveals ancestral eumetazoan gene repertoire and genomic organization
|journal=[[Science (journal)|Science]]
|volume=317
|issue=5834
|pages=86–94
|doi=10.1126/science.1139158
|pmid=17615350
|bibcode=2007Sci...317...86P}}</ref>

An analysis of the [[Homoscleromorpha|homoscleromorph sponge]] ''Oscarella carmela'' also suggests that the last common ancestor of sponges and the eumetazoan animals was more complex than previously assumed.<ref>{{cite journal | coauthors = Wang, Xiujuan; Lavrov Dennis V. | date = 27 October 2006 | title = Mitochondrial Genome of the homoscleromorph Oscarella carmela ([[Porifera]], [[Demospongiae]]) Reveals Unexpected Complexity in the Common Ancestor of Sponges and Other Animals | journal = Molecular Biology and Evolution | volume = 24 | issue = 2 | pages = 363–373| doi = 10.1093/molbev/msl167 | url = http://mbe.oxfordjournals.org/cgi/content/abstract/24/2/363 | author = Wang, X. | pmid = 17090697 | first1 = X | first2 = DV}}</ref>

Other model organisms belonging to the animal kingdom include the house mouse (''[[Mus musculus]]'') and zebrafish (''[[Danio rerio]]'').

[[File:Carolus Linnaeus (cleaned up version).jpg|thumb|upright|[[Carolus Linnaeus]], known as the father of modern [[taxonomy]]]]

==History of classification==
[[Aristotle]] divided the living world between animals and plants, and this was followed by [[Carolus Linnaeus]] (Carl von Linné), in the first hierarchical classification.<ref name="Linn1758">{{cite book|last=Linnaeus |first=Carolus |authorlink=Carl Linnaeus |title=Systema naturae per regna tria naturae :secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. |publisher=Holmiae (Laurentii Salvii) |year=1758 |url=http://www.biodiversitylibrary.org/bibliography/542 |accessdate=22 September 2008 |language=Latin |edition=[[10th edition of Systema Naturae|10th edition]]| archiveurl= http://web.archive.org/web/20081010032456/http://www.biodiversitylibrary.org/bibliography/542| archivedate= 10 October 2008 <!--DASHBot-->| deadurl= no}}</ref> Since then biologists have begun emphasizing evolutionary relationships, and so these groups have been restricted somewhat. For instance, microscopic [[protozoa]] were originally considered animals because they move, but are now treated separately.

In Linnaeus's original scheme, the animals were one of three kingdoms, divided into the classes of [[Vermes in the 10th edition of Systema Naturae|Vermes]], [[Insecta in the 10th edition of Systema Naturae|Insect]]a, [[Pisces in the 10th edition of Systema Naturae|Pisces]], [[Amphibia in the 10th edition of Systema Naturae|Amphibia]], [[Aves in the 10th edition of Systema Naturae|Aves]], and [[Mammalia in the 10th edition of Systema Naturae|Mammalia]]. Since then the last four have all been subsumed into a single phylum, the [[chordate|Chordata]], whereas the various other forms have been separated out. The above lists represent our current understanding of the group, though there is some variation from source to source.

==Ethics==
{{Expand section|date=October 2012}}

{{Main|Animal rights}}

Animals are regarded by philosophers like [[Peter Singer]] as [[ethics|ethical]] subjects. Animal rights movements have worked to promote animal welfare.

==See also==
{{Portal|Animals}}
{{Wikipedia books}}
{{Wikispecies| Animalia}}
* [[Ethology]]
* [[Animal colouration]]
* [[Animal rights]]
* [[Fauna]]
* [[List of animal names]]
* [[List of animals by number of neurons]]
* [[Lists of animals]]
<div style="clear:both"></div>

==References==
{{Reflist|colwidth=25em}}

===Bibliography===
* Klaus Nielsen. ''Animal Evolution: Interrelationships of the Living Phyla'' (2nd edition). [[Oxford University Press]], 2001.
* Knut Schmidt-Nielsen. ''Animal Physiology: Adaptation and Environment''. (5th edition). [[Cambridge University Press]], 1997.

==External links==
{{Sister project links|Animalia}}
* {{eol|1}}
* [http://tolweb.org/ Tree of Life Project]
* [http://animaldiversity.ummz.umich.edu/site/index.html Animal Diversity Web] – [[University of Michigan]]'s database of animals, showing [[taxonomic classification]], images, and other information.
* [http://www.arkive.org ARKive] – multimedia database of worldwide endangered/protected species and common species of UK.
* [http://www.sciam.com/article.cfm?chanID=sa006&articleID=000DC8B8-EA15-137C-AA1583414B7F0000 Scientific American Magazine (December 2005 Issue) – Getting a Leg Up on Land] About the evolution of four-limbed animals from fish.

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[[lij:Animalia]]
[[li:Bieste]]
[[ln:Nyama]]
[[jbo:danlu]]
[[lmo:Bestia]]
[[hu:Állatok]]
[[mk:Животно]]
[[mg:Biby]]
[[ml:ജന്തു]]
[[mt:Annimal]]
[[mr:प्राणी]]
[[xmf:ჩხოლარეფი]]
[[mzn:حیوون]]
[[ms:Haiwan]]
[[mwl:Animal]]
[[mn:Амьтан]]
[[my:တိရစ္ဆာန်]]
[[nah:Yōlcatl]]
[[fj:Manumanu]]
[[nl:Dierenrijk]]
[[cr:ᐱᓯᐢᑭᐤ]]
[[ne:जनावर]]
[[ja:動物]]
[[nap:Animali]]
[[frr:Diarten]]
[[no:Dyr]]
[[nn:Dyr]]
[[nrm:Animâ]]
[[nov:Animalia]]
[[oc:Animalia]]
[[mhr:Янлык]]
[[om:Binensotta]]
[[uz:Xayvonlar]]
[[pnb:جاندار]]
[[ps:ژوي]]
[[koi:Пода]]
[[pms:Animalia]]
[[tpi:Abus]]
[[nds:Deert]]
[[pl:Zwierzęta]]
[[pt:Animalia]]
[[ksh:Dier]]
[[ro:Regnul Animalia]]
[[qu:Uywa]]
[[rue:Быдла]]
[[ru:Животные]]
[[sah:Харамай]]
[[sa:जन्तवः]]
[[sco:Ainimal]]
[[sq:Kafsha]]
[[scn:Armali]]
[[si:සත්තු]]
[[simple:Animal]]
[[sk:Živočíchy]]
[[sl:Živali]]
[[so:Xayawaan]]
[[srn:Meti]]
[[sr:Животиње]]
[[sh:Životinje]]
[[su:Sato]]
[[fi:Eläinkunta]]
[[sv:Djur]]
[[tl:Hayop]]
[[ta:விலங்கு]]
[[roa-tara:Regnum Animalia]]
[[tt:Хайваннар]]
[[te:జంతువు]]
[[th:สัตว์]]
[[ti:እንስሳ]]
[[tg:Ҳайвон]]
[[to:Monumanu]]
[[chr:ᎦᏂᏝᎢ]]
[[chy:Hova]]
[[ve:Phukha]]
[[tr:Hayvanlar]]
[[uk:Тварини]]
[[ur:جانور]]
[[ug:ھايۋانات]]
[[vi:Động vật]]
[[fiu-vro:Eläjäq]]
[[wa:Biesse]]
[[zh-classical:動物]]
[[vls:Bêesten (ryk)]]
[[war:Hayop]]
[[wo:Dundat yi]]
[[yi:בעלי חיים]]
[[yo:Ẹranko]]
[[zh-yue:動物]]
[[zea:Beêsten]]
[[bat-smg:Gīvūnā]]
[[zh:动物]]

Revision as of 01:39, 7 December 2012

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