Cladonychiidae

Cladonychiidae
	Briggsus, Oregon
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Opiliones
Infraorder:	Insidiatores
Superfamily:	Travunioidea
Family:	Cladonychiidae; Hadži, 1935
Synonyms
	Erebromastridae

The Cladonychiidae are a small family of harvestman with about 33 described species, within the suborder Laniatores.

Description

Members of this family range from less than two to about four millimeters in body length, with robust, spined pedipalps and rather short legs, although the second pair can be as long as two centimeter. Most Cladonychiidae are reddish brown to dark brown, but cave-dwelling species are pale yellow. Not all species have eyes.^[1]

Distribution

The members of Cladonychiidae are found in Southern Europe and the United States.^[2]

Fossils

Proholoscotolemon was recently found in Baltic amber.

Relationships

The genera Peltonychia, Holoscotolemon, Erebomaster, Theromaster, Speleonychia, Briggsus, and Isolachus from the family Travuniidae and the former family Pentanychidae have been transferred to Cladonychiidae. This left Travuniidae with three genera from the Balkan region of Europe, Travunia, Trojanella, and Dinaria.^[2]

Name

The name of the former type genus Cladonychium (now synonymized with Erebomaster) is derived from Ancient Greek "branched claw".^[1]

Species

These genera and species belong to the family Cladonychiidae:^[2]^[3]^[4]

(For Arbasus Roewer, 1935 see Buemarinoidae Karaman, 2019)

genus Briggsus Özdikmen & Demir, 2008
- species Briggsus bilobatus (Briggs, 1971)
- species Briggsus clavatus (Briggs, 1971)
- species Briggsus flavescens (Briggs, 1971)
- species Briggsus hamatus (Briggs, 1971)
- species Briggsus pacificus (Briggs, 1971)
genus Buemarinoa Roewer, 1956
- species Buemarinoa patrizii Roewer, 1956
genus Erebomaster Cope, 1872
- species Erebomaster acanthinus (Crosby & Bishop, 1924)
- species Erebomaster flavescens Cope, 1872
  - subspecies Erebomaster flavescens flavescens Cope, 1872
  - subspecies Erebomaster flavescens coecum (Packard, 1888) — Carter cave
- species Erebomaster weyerensis (Packard, 1888)
genus Holoscotolemon Roewer, 1915
- species Holoscotolemon franzinii Tedeschi & Sciaky, 1994
- species Holoscotolemon jaqueti (Corti, 1905)
- species Holoscotolemon lessiniense Martens, 1978
- species Holoscotolemon lessiniensis Martens, 1978
- species Holoscotolemon monzinii Tedeschi & Sciaky, 1994
- species Holoscotolemon naturae Tedeschi & Sciaky, 1994
- species Holoscotolemon oreophilus Martens, 1978
- species Holoscotolemon querilhaci (Lucas, 1864)
- species Holoscotolemon unicolor Roewer, 1915
genus Isolachus Briggs, 1971
- species Isolachus spinosus Briggs, 1971
genus Peltonychia Roewer, 1935
- species Peltonychia clavigera (Simon, 1879)
- species Peltonychia gabria Roewer, 1935
- species Peltonychia leprieuri (Lucas, 1861)
- species Peltonychia leprieurii
- species Peltonychia navarica (Simon, 1879)
- species Peltonychia piochardi (Simon, 1872)
- species Peltonychia postumicola (Roewer, 1935)
- species Peltonychia sarea (Roewer, 1935)
- species Peltonychia tenuis Roewer, 1935
genus Speleonychia Briggs, 1974
- species Speleonychia sengeri Briggs, 1974
genus Theromaster Briggs, 1969
- species Theromaster archeri (Goodnight & Goodnight, 1942)
- species Theromaster brunneus (Banks, 1902)
genus † Proholoscotolemon Ubick & Dunlop, 2005
- species † Proholoscotolemon nemastomoides (Koch & Berendt, 1854)

Footnotes

^ ^a ^b Briggs, Thomas S. & Ubick, Darrell (2007): Cladonychiidae Hadži, 1935. In: Pinto-da-Rocha et al. 2007: 179ff
^ ^a ^b ^c Derkarabetian, Shahan; Starrett, James; Tsurusaki, Nobuo; Ubick, Darrell; et al. (2018). "A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements". ZooKeys (760): 1–36. doi:10.3897/zookeys.760.24937. PMC 5986891. PMID 29872361.
^ Kury, Adriano B. (2018). "Familial nomina in harvestmen (Arachnida, Opiliones)". Bionomina. 13 (1): 1–27. doi:10.11646/bionomina.13.1.1. S2CID 92517624.
^ Kury, Adriano Brilhante (2013). Zhang, Zhi-Qiang (ed.). "Order Opiliones Sundevall, 1833. In: Zhang Z-Q (ed) Animal biodiversity: An outline of higher-level classif. and survey of taxonomic richness (Addenda 2013)". Zootaxa. 3703 (1). doi:10.11646/zootaxa.3703.1.7. ISBN 978-1-86977-849-1.

References

Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds.) (2007): Harvestmen - The Biology of Opiliones. Harvard University Press ISBN 0-674-02343-9

[pinto-179ff-1] Briggs, Thomas S. & Ubick, Darrell (2007): Cladonychiidae Hadži, 1935. In: Pinto-da-Rocha et al. 2007: 179ff

[Derkarabetian2018-2] Derkarabetian, Shahan; Starrett, James; Tsurusaki, Nobuo; Ubick, Darrell; et al. (2018). "A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements". ZooKeys (760): 1–36. doi:10.3897/zookeys.760.24937. PMC 5986891. PMID 29872361.

[Kury2018-3] Kury, Adriano B. (2018). "Familial nomina in harvestmen (Arachnida, Opiliones)". Bionomina. 13 (1): 1–27. doi:10.11646/bionomina.13.1.1. S2CID 92517624.

[Kury2013-4] Kury, Adriano Brilhante (2013). Zhang, Zhi-Qiang (ed.). "Order Opiliones Sundevall, 1833. In: Zhang Z-Q (ed) Animal biodiversity: An outline of higher-level classif. and survey of taxonomic richness (Addenda 2013)". Zootaxa. 3703 (1). doi:10.11646/zootaxa.3703.1.7. ISBN 978-1-86977-849-1.

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