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{{Taxobox
| color = pink
| name = Turbonillinae
| image =
| image_width = 240px
| image_caption =
| regnum = [[Animal]]ia
| phylum = [[Mollusca]]
| classis = [[Gastropoda]]
| ordo = [[Heterostropha]]
| superfamilia = [[Pyramidelloidea]]
| familia = [[Pyramidellidae]]
| subfamilia = '''Turbonillinae'''
| subfamilia_authority = H.G. Bronn, 1849
| subdivision_ranks = Genera
| subdivision = See text.
| synonyms = Chemniziidae <small>Stoliczka, 1868</small>
}}


'''Turbonillinae''' is a [[subfamily]] of mostly minute [[parasite|parasitic]] sea [[snail]]s, [[marine (ocean)|marine]] [[gastropod]] [[mollusc]]s in the family [[Pyramidellidae]], the '''pyrams''' and their allies. This is one of eleven recognised subfamilies in the family [[Pyramidellidae]] (according to the taxonomy of Ponder & Lindberg, 1997).

However, in the [[Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)|taxonomy of Bouchet & Rocroi]] (2005), this subfamily also comprises the subfamilies Cingulininae and Eulimellinae, that have been downgraded to the rank of [[tribe (biology)|tribe]] Cingulinini and Eulimellini
*Subfamily [[Turbonillinae]] <small> Bronn, 1849 </small>
**Tribe Turbonillini <small> Bronn, 1849 </small>
**Tribe Cingulinini <small> Saurin, 1958 </small> (formerly subfamily [[Cingulininae]])
**Tribe Eulimellini <small> Saurin, 1958 </small> (formerly subfamily [[Eulimellinae]])

According to Schander, Van Aartsen & Corgan (1999) there are 27 Genera within the [[Turbonillinae]].

==Genera within the subfamily Turbonillinae==
* ''[[Turbonilla]]'' <small> Risso, 1826 </small> (Type genus)
* ''[[Amamimormula]]'' <small> Kuroda, 1928</small>
* ''[[Asmunda]]'' <small> Dall & Bartsch, 1904</small>
* ''[[Bartschella (mollusk)|Bartschella]]'' <small> Iredale, 1916</small>
* ''[[Baldra]]'' <small> Dall & Bartsch, 1904</small>
* ''[[Cylindroturbonilla]]'' <small> Nordsieck, 1972</small>
* ''[[Exesilla]]'' <small> Laseron, 1959</small>
* ''[[Gispyrella]]'' <small> Laws, 1937</small>
* ''[[Graciliturbonilla]]'' <small> Nordsieck, 1972</small>
* ''[[Houbrickia]]'' <small> Wise, 1996</small>
* ''[[Lancea (mollusk)|Lancea]]'' <small> Peace, 1868</small>
* ''[[Magniturbonilla]]'' <small> Nordsieck, 1972</small>
* ''[[Mormula]]'' <small>A. Adams, 1863</small>
* ''[[Nisiturris]]'' <small> Dall & Bartsch, 1906</small>
* ''[[Paramormula]]'' <small> Nomura, 1939</small>
* ''[[Paraturbonilla]]'' <small> Boettger, 1906</small>
* ''[[Planpyrgiscus]]'' <small> Laws, 1937</small>
* ''[[Pselliogyra]]'' <small> Dall & Bartsch, 1909</small>
* ''[[Pyrgiscilla]]'' <small> Laws, 1937</small>
* ''[[Pyrgiscus]]'' <small> Philippi, 1841</small>
* ''[[Pyrgolampros]]'' <small> Sacco, 1892</small>
* ''[[Pyrgolidium]]'' <small> Monterosato, 1884</small>
* ''[[Striarcana]]'' <small> Laws, 1937</small>
* ''[[Sulcoturbonilla]]'' <small> Sacco, 1892</small>
* ''[[Tereoturbonilla]]'' <small> Eames, 1951</small>
* ''[[Zaphella]]'' <small> Laseron, 1959</small>

==Problematic genera within the family [[Turbonillidae]]==
The following genera are currently difficult to place within the family [[Turbonillidae]] (=[[Turbonillinae]] + [[Cingulininae]] + [[Eulimellinae]])
* ''[[Atomiscala]]'' <small> DeBoury, 1909 </small>
* ''[[Hamarilla]]'' <small> Eames & Wilkins, 1957 </small>
* ''[[Pseudographis]]'' <small> Mifsud, 1998 </small>
* ''[[Rissopsetia]]'' <small> Dell, 1956 </small>

==Distribution==
This family is found worldwide.

==Shell description==
The shell of these snails has a blunt, [[heterostrophic]] [[protoconch]], which is pointed sideways. The shell of most species are rather high and slender.

The texture of these shells is usually ribbed [[sculpture (mollusc)|sculptured]] in various forms and often also have more or less prominent spirals. Their color is mostly white, cream or yellowish, sometimes with red or brown lines.

The adult shell, the [[whorl (mollusk)|teleoconch]] is [[wiktionary:dextral|dextrally]] coiled, but the [[larva]]l shells are [[wiktionary:sinistral|sinistral]]. This results in a sinistrally coiled protoconch. The opening of the shell, the [[aperture (mollusc)|aperture]] is closed by a lid, a so called [[Operculum (gastropod)|operculum]].

==Life habits==
The Turbonillinae are [[ectoparasite]]s, feeding mainly on other molluscs and on annelid [[worm]]s.

They do not have a [[radula]]. Instead their long [[proboscis]] is used to pierce the skin of its prey and suck up its fluids and soft tissues. The eyes on the grooved tentacles are situated toward the base of the tentacles. Between the head and the foot, a lobed process called the [[mentum]] ( = thin projection) is visible.

These molluscs are [[hermaphrodite]]s, laying eggs in jelly-like masses on the shell of its host.


==References==
* {{cite journal
| author = Bouchet, P. & Rocroi, J.-P.
| year = 2005
| date=
| title = Classification and Nomenclator of Gastropod Families
| journal = Malacologia
| volume =47
| issue = 1-2
| pages = 1-397
| issn =
| doi =
}}
* {{cite journal
| author = Ponder, W.F. & Lindberg, D. R.
| year = 1997
| date=
| title = Towards a phylogeny of gastropod molluscs: an analysis using morphological characters
| journal = Zoological Journal of the Linnean Society
| volume =119
| issue =
| pages = 88-265
| issn =
| doi =
}}
* {{cite journal
| author = Schander, C., van Aartsen, J. J., Corgan, J. C.
| year = 1999
| date=
| title = Families and genera of the Pyramidelloidea (Mollusca: Gastropoda)
| journal = Bollettino Malacologico
| volume =34
| issue = 9-12
| pages = 145-166
| issn =
| doi =
}}

* {{listdev}}


[[Category:Pyramidellidae]]
[[Category:Pyramidellidae]]

Revision as of 13:33, 26 January 2008