|This article needs additional citations for verification. (February 2014)|
The pennaceous feather is a type of feather present in most modern birds and in some species of maniraptoriform dinosaurs. A pennaceous feather has a stalk or quill. Its basal part, called a calamus, is embedded in the skin. The calamus is hollow and has pith formed from the dry remains of the feather pulp, and the calamus opens below by an inferior umbilicus and above by a superior umbilicus. The stalk above the calamus is a solid rachis having an umbilical groove on its underside. Pennaceous feathers have a rachis with vanes or vaxillum spreading to either side. These vanes are composed of a high number of flattened barbs, that are connected to one another with barbules.
Pennaceous feathers on the forelimb or wing are often attached strongly due to stresses related to flight or other activities. This strong attachment is accomplished by ligaments under the skin, which in some birds and feathered dinosaurs results in raised bumps or marks along the rear forelimb bone (ulna). These bumps, called quill knobs (ulnar papillae) are often used as an indirect indication of strongly-attached forelimb feathers in fossil species, and can also indirectly indicate the number of secondary regimes in a given specimen.
- A Kovalev, AE Filippov, SN Gorb (2014) Unzipping bird feathers. J. R. Soc. Interface 11 (92) 20130988. doi:10.1098/rsif.2013.0988
- Turner, A.H.; Makovicky, P.J.; Norell, M.A. (2007). "Feather quill knobs in the dinosaur Velociraptor". Science 317 (5845): 1721. Bibcode:2007Sci...317.1721T. doi:10.1126/science.1145076. PMID 17885130.
- Zelenitsky, D. K.; Therrien, F.; Erickson, G. M.; Debuhr, C. L.; Kobayashi, Y.; Eberth, D. A.; Hadfield, F. (2012). "Feathered Non-Avian Dinosaurs from North America Provide Insight into Wing Origins". Science 338 (6106): 510–514. doi:10.1126/science.1225376. PMID 23112330.
|This article about ornithology is a stub. You can help Wikipedia by expanding it.|