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'''''Worthenella''''' is a genus of enigmatic [[arthropod]] from the [[Burgess Shale]]. It known from a single specimen described initially as an [[annelid]] by [[Charles Doolittle Walcott]] in 1911. The body is elongate and [[myriapod]]-like with a head and at least 46 body segments. The head bears poorly preserved appendages, while filamentous branched structures appear to run along the underside of the first 34 trunk segments, with the posterior 8 suggested to bare longer appendages.<ref>{{Cite web |title=''Worthenella cambria'' |url=https://burgess-shale.rom.on.ca/fossils/worthenella-cambria/ |access-date=2023-01-13 |website=The Burgess Shale |publisher=Royal Ontario Museum |language=en-US}}</ref> In 2013, David Legg placed it in the family [[Kootenichelidae]] as a sister to ''[[Kootenichela]]'' , based on the supposed presence of antenniform head appendages.<ref>{{Cite journal | doi = 10.1666/12-112.1| title = Multi-Segmented Arthropods from the Middle Cambrian of British Columbia (Canada)| journal = Journal of Paleontology| volume = 87| issue = 3| pages = 493–501| year = 2013| last1 = Legg | first1 = David | bibcode = 2013JPal...87..493L| s2cid = 86725173}}</ref> However, this position was questioned in a later study, which argued that the supposed annteniform appendages were actually [[taphonomic]] artefacts.<ref>{{Cite journal |last1=Lamsdell |first1=James C. |last2=Stein |first2=Martin |last3=Selden |first3=Paul A. |date=September 2013 |title=Kodymirus and the case for convergence of raptorial appendages in Cambrian arthropods |url=http://link.springer.com/10.1007/s00114-013-1081-y |journal=Naturwissenschaften |language=en |volume=100 |issue=9 |pages=811–825 |doi=10.1007/s00114-013-1081-y |pmid=23893175 |bibcode=2013NW....100..811L |s2cid=253635469 |issn=0028-1042}}</ref>
'''''Worthenella''''' is a genus of enigmatic [[arthropod]] from the [[Burgess Shale]]. It known from a single specimen described initially as an [[annelid]] by [[Charles Doolittle Walcott]] in 1911. The body is elongate and [[myriapod]]-like with a head and at least 46 body segments. The head bears poorly preserved appendages, while filamentous branched structures appear to run along the underside of the first 34 trunk segments, with the posterior 8 suggested to have had longer appendages.<ref>{{Cite web |title=''Worthenella cambria'' |url=https://burgess-shale.rom.on.ca/fossils/worthenella-cambria/ |access-date=2023-01-13 |website=The Burgess Shale |publisher=Royal Ontario Museum |language=en-US}}</ref> In 2013, David Legg placed it in the family [[Kootenichelidae]] as a sister to ''[[Kootenichela]]'' , based on the supposed presence of antenniform head appendages.<ref>{{Cite journal | doi = 10.1666/12-112.1| title = Multi-Segmented Arthropods from the Middle Cambrian of British Columbia (Canada)| journal = Journal of Paleontology| volume = 87| issue = 3| pages = 493–501| year = 2013| last1 = Legg | first1 = David | bibcode = 2013JPal...87..493L| s2cid = 86725173}}</ref> However, this position was questioned in a later study, which argued that the supposed annteniform appendages were actually [[taphonomic]] artefacts.<ref>{{Cite journal |last1=Lamsdell |first1=James C. |last2=Stein |first2=Martin |last3=Selden |first3=Paul A. |date=September 2013 |title=Kodymirus and the case for convergence of raptorial appendages in Cambrian arthropods |url=http://link.springer.com/10.1007/s00114-013-1081-y |journal=Naturwissenschaften |language=en |volume=100 |issue=9 |pages=811–825 |doi=10.1007/s00114-013-1081-y |pmid=23893175 |bibcode=2013NW....100..811L |s2cid=253635469 |issn=0028-1042}}</ref>


==References==
==References==

Revision as of 03:32, 25 November 2023

Worthenella
Temporal range: Burgess Shale
Type specimen of W. cambria in shale
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: incertae sedis
Family: Kootenichelidae
Genus: Worthenella
Walcott, 1911[1]
Type species
Worthenella cambria
Walcott, 1911[1]

Worthenella is a genus of enigmatic arthropod from the Burgess Shale. It known from a single specimen described initially as an annelid by Charles Doolittle Walcott in 1911. The body is elongate and myriapod-like with a head and at least 46 body segments. The head bears poorly preserved appendages, while filamentous branched structures appear to run along the underside of the first 34 trunk segments, with the posterior 8 suggested to have had longer appendages.[2] In 2013, David Legg placed it in the family Kootenichelidae as a sister to Kootenichela , based on the supposed presence of antenniform head appendages.[3] However, this position was questioned in a later study, which argued that the supposed annteniform appendages were actually taphonomic artefacts.[4]

References

  1. ^ a b Walcott, Charles D. (1911). "Cambrian Geology and Paleontology II. Middle Cambrian annelids". Smithsonian Miscellaneous Collections. 57 (5): 125–127. hdl:10088/34820Pl. 22, Fig. 2. {{cite journal}}: External link in |postscript= (help)CS1 maint: postscript (link)
  2. ^ "Worthenella cambria". The Burgess Shale. Royal Ontario Museum. Retrieved 2023-01-13.
  3. ^ Legg, David (2013). "Multi-Segmented Arthropods from the Middle Cambrian of British Columbia (Canada)". Journal of Paleontology. 87 (3): 493–501. Bibcode:2013JPal...87..493L. doi:10.1666/12-112.1. S2CID 86725173.
  4. ^ Lamsdell, James C.; Stein, Martin; Selden, Paul A. (September 2013). "Kodymirus and the case for convergence of raptorial appendages in Cambrian arthropods". Naturwissenschaften. 100 (9): 811–825. Bibcode:2013NW....100..811L. doi:10.1007/s00114-013-1081-y. ISSN 0028-1042. PMID 23893175. S2CID 253635469.