Bembridae
| Bembridae | |
|---|---|
| |
| Bembradium roseum | |
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Order: | Scorpaeniformes |
| Suborder: | Platycephaloidei |
| Family: | Bembridae Kaup, 1873[1] |
| Genera | |
|
see text | |
Bembridae, the deep-water flatheads, are a family of bottom-dwelling ray-finned fishes. They are found in the Indian and western Pacific Oceans.
Taxonomy
Bembridae was first proposed as a family in 1873 by the German zoologist Johann Jakob Kaup.[1] The 5th edition of Fishes of the World classifies this family within the suborder Platycephaloidei in the order Scorpaeniformes.[2] Other authorities differ and do not consider the Scorpaeniformes to be a valid order because the Perciformes is not monophyletic without the taxa within the Scorpaeniformes being included within it. These authorities consider the Bembridae to belong to the suborder Platycephaloidei, along with the families Parabembridae, Hoplichthyidae, Platycephalidae and Plectrogeniidae within the Perciformes.[3] This family is thought to be more primitive than their close relatives, the true flatheads. Despite the common name, their heads are only slightly flattened and have spiny ridges.[4]
Genera
Bembridae contains 5 recognised genera:[2]
- Bambradon D.S. Jordan & R.E. Richardson, 1908[5]
- Bembradium Gilbert, 1905[6]
- Bembras Cuvier, 1829[5]
- Brachybembras Fowler. 1938[5]
- Parabembras Bleeker, 1874[5]
Other authorities classify the genus Parabembras within the monotypic family Parabembridae and place Bembradium in the family Plectrogeniidae.[3]
Etymology
Bembridae derives its name from its type genus Bembras which was coined by the French zoologist Georges Cuvier in 1829 but he did not explain its etymology. However, it is thought that it may come from an ancient Greek word for some sort of small fish, such as anchovy, sprat or smelt. which at least dates as far back as Aristotle. Cuvier applied this type of name to other genera he put forward, such as Synodontis, Salanx or Premnas.[7]
Characteristics
Benbridae deepwater flatheads have large, flattened heads with elongated subcylindrical bodies which have not spiny scutes. The eyes are moderately large without an iris lapit or ocular flaps. The infraorbital bones form a clear, robust ridge, joining the uppermost preopercular spine and has between 4 and 6 spines on it. The spines on the head are short to moderate with between 8 and 11 above the eye. There are two separate dorsal fins, the first contains between 6 and 12 spines and the second between 8 and 12 soft rays while there are 3 spines in the anal fin. There are no separate rays in the pectoral fins. Although the colour is typically red red, orange, or light brown; some species have green, dark green or blackish spotting or blotching on their body and fins and the anal fin may be almost black.[2][8] The deepwater flatheads range in size from Bembradium roseum, which has a maximum published standard length of 11 cm (4.3 in), to the bigscale flathead (Bembras macrolepis), which reaches a standard length of 27.3 cm (10.7 in).[9]
Distribution and habitat
Bembridae deep-water flatheads have an Indo-Pacific distribution and are mainly found in the South China Sea, the Gulf of Aden, and the waters near Southern Indonesia and Australia.[10] These are benthic fishes of the continental shelf at depths of 80 to 581 m (262 to 1,906 ft).[8]
References
- ^ a b Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014). "Family-group names of Recent fishes". Zootaxa. 3882 (2): 001–230. doi:10.11646/zootaxa.3882.1.1. PMID 25543675.
- ^ a b c J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). Fishes of the World (5th ed.). Wiley. pp. 467–495. ISBN 978-1-118-34233-6. Archived from the original on 2019-04-08. Retrieved 2022-06-30.
- ^ a b Ricardo Betancur-R; Edward O. Wiley; Gloria Arratia; et al. (2017). "Phylogenetic classification of bony fishes". BMC Evolutionary Biology. 17 (162): 162. doi:10.1186/s12862-017-0958-3. PMC 5501477. PMID 28683774.
- ^ Eschmeyer, William N. (1998). Paxton, J.R.; Eschmeyer, W.N. (eds.). Encyclopedia of Fishes. San Diego: Academic Press. p. 177. ISBN 978-0-12-547665-2.
- ^ a b c d Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Genera in the family Bemridae". Catalog of Fishes. California Academy of Sciences. Retrieved 30 June 2022.
- ^ Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Genera in the family Pectrogeniidae". Catalog of Fishes. California Academy of Sciences. Retrieved 30 June 2022.
- ^ Christopher Scharpf & Kenneth J. Lazara, eds. (7 December 2021). "Order Perciformes (Part 11): Suborder Platycephaloidei: Families Bembridae, Parabembridae, Hoplichthyidae, Platycephalidae and Plectrogeniidae". The ETYFish Project Fish Name Etymology Database. Christopher Scharpf and Kenneth J. Lazara. Retrieved 26 May 2022.
- ^ a b S.W. Poss (1999). "Bembridae Deepwater flatheads". In Carpenter, K.E.; Niem, V.H. (eds.). FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Volume 4. Bony fishes part 2 (Mugilidae to Carangidae). FAO, Rome. pp. 2383–2384. ISBN 9251043019.
- ^ Froese, Rainer; Pauly, Daniel (eds.). "Family Bembridae". FishBase. February 2022 version.
- ^ Imamura, Hisashi; Knapp, Leslie W. (1997). "A new species of deepwater flathead, Bembras adenensis (Scorpaeniformes: Bembridae) from the western Indian Ocean". Ichthyological Research. 44 (1): 9. doi:10.1007/bf02672753. hdl:1834/955. ISSN 1341-8998.