Jump to content

Azhdarchidae

From Wikipedia, the free encyclopedia

This is an old revision of this page, as edited by Chaoyangopterus (talk | contribs) at 19:06, 26 October 2015. The present address (URL) is a permanent link to this revision, which may differ significantly from the current revision.

Azhdarchids
Temporal range: Late Cretaceous, 108–66 Ma Possibly earlier
Reconstructed skeleton of Quetzalcoatlus northropi
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: †Pterosauria
Suborder: †Pterodactyloidea
Clade: †Azhdarchiformes
Family: †Azhdarchidae
Nesov, 1984
Type species
†Azhdarcho lancicollis
Nesov, 1984
Genera

See text

Synonyms

"Titanopterygiidae"
Padian, 1984 (preoccupied)

Azhdarchidae (from اژدرها (Aždarha), the Persian word for dragon) is a family of pterosaurs known primarily from the late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the early Cretaceous as well (late Berriasian age, about 140Ma ago).[1] Azhdarchids included some of the largest known flying animals of all time. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and "Titanopteryx" (now known as Arambourgiania). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil reccord, but recent studies indicate a wealth in pterosaurian faunas, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms.[2] Some taxa like Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.[3][4][5]

Description

Artist's reconstruction of foraging Quetzalcoatlus northropi

Azhdarchids are characterized by their long legs and extremely long necks, made up of elongated neck vertebrae which are round in cross section. Most species of azhdarchids are still known mainly from their distinctive neck bones and not much else. The few azhdarchids that are known from reasonably good skeletons include Zhejiangopterus and Quetzalcoatlus. Azhdarchids are also distinguished by their relatively large heads and long, spear-like jaws. It had been suggested azhdarchids were skimmers,[6][7] but further research has cast doubt on this idea, demonstrating that azhdarchids lacked the necessary adaptations for a skim-feeding lifestyle, and that they may have led a more terrestrial existence similar to modern storks and ground hornbills.[8][9][10][11][12]

Systematics

Azhdarchids were originally classified as close relatives of Pteranodon due to their long, toothless beaks. Others have suggested they were more closely related to the toothy Ctenochasmatids (which include filter-feeders like Ctenochasma and Pterodaustro). Currently it is widely agreed that azhdarchids were closely related to pterosaurs such as Tupuxuara and Tapejara.

Taxonomy

Classification after Unwin 2006, except where noted.[13]

Reconstructed feeding posture of an azhdarchid with sagittally aligned limbs.

References

  1. ^ Dyke, G., Benton, M., Posmosanu, E. and Naish, D. (2010). "Early Cretaceous (Berriasian) birds and pterosaurs from the Cornet bauxite mine, Romania." Palaeontology, published online before print 15 September 2010. doi:10.1111/j.1475-4983.2010.00997.x
  2. ^ Agnolin, Federico L. and Varricchio, David (2012). "Systematic reinterpretation of Piksi barbarulna Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird" (PDF). Geodiversitas. 34 (4): 883–894. doi:10.5252/g2012n4a10.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  3. ^ Carroll, N. REASSIGNMENT OF MONTANAZHDARCHO MINOR AS A NON-AZHDARCHID MEMBER OF THE AZHDARCHOIDEA, SVP 2015
  4. ^ a b Andres, B.; Myers, T. S. (2013). "Lone Star Pterosaurs". Earth and Environmental Science Transactions of the Royal Society of Edinburgh: 1. doi:10.1017/S1755691013000303.
  5. ^ Wilton, Mark P. (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. ISBN 0691150613.
  6. ^ Nesov, L. A. (1984). "Upper Cretaceous pterosaurs and birds from Central Asia". Paleontologicheskii Zhurnal. 1984 (1): 47–57.
  7. ^ Kellner, A. W. A.; Langston, W. (1996). "Cranial remains of Quetzalcoatlus (Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas". Journal of Vertebrate Paleontology. 16 (2): 222–231. doi:10.1080/02724634.1996.10011310.
  8. ^ Chatterjee, S.; Templin, R. J. (2004). "Posture, locomotion, and paleoecology of pterosaurs". Geological Society of America Special Publication. 376: 1–64. doi:10.1130/0-8137-2376-0.1.
  9. ^ Ősi, A.; Weishampel, D.B.; Jianu, C.M. (2005). "First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary". Acta Palaeontologica Polonica. 50 (4): 777–787.
  10. ^ Humphries, S.; Bonser, R.H.C.; Witton, M.P.; Martill, D.M. (2007). "Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method" (PDF). PLoS Biology. 5 (8): e204. doi:10.1371/journal.pbio.0050204.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  11. ^ Witton, Mark P.; Naish, Darren; McClain, Craig R. (28 May 2008). "A Reappraisal of Azhdarchid Pterosaur Functional Morphology and Paleoecology". PLoS ONE. 3 (5): e2271. doi:10.1371/journal.pone.0002271. PMC 2386974. PMID 18509539.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  12. ^ "Pterosaurs".
  13. ^ Unwin, David M. (2006). The Pterosaurs: From Deep Time. New York: Pi Press. p. 273. ISBN 0-13-146308-X.
  14. ^ Ibrahim, N.; Unwin, D.M.; Martill, D.M.; Baidder, L.; Zouhri, S. (2010). Farke, Andrew Allen (ed.). "A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco". PLoS ONE. 5 (5): e10875. doi:10.1371/journal.pone.0010875. PMC 2877115. PMID 20520782.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  15. ^ Averianov, A.O. (2007). "New records of azhdarchids (Pterosauria, Azhdarchidae) from the late Cretaceous of Russia, Kazakhstan, and Central Asia". Paleontological Journal. 41 (2): 189–197. doi:10.1134/S0031030107020098.
  16. ^ a b Averianov, A.O. (2010). "The osteology of Azhdarcho lancicollis Nessov, 1984 (Pterosauria, Azhdarchidae) from the Late Cretaceous of Uzbekistan." Proceedings of the Zoological Institute of the Russian Academy of Sciences, 314(3): 246-317.
  17. ^ Vremir, M. T. S.; Kellner, A. W. A.; Naish, D.; Dyke, G. J. (2013). Viriot, Laurent (ed.). "A New Azhdarchid Pterosaur from the Late Cretaceous of the Transylvanian Basin, Romania: Implications for Azhdarchid Diversity and Distribution". PLoS ONE. 8: e54268. doi:10.1371/journal.pone.0054268.{{cite journal}}: CS1 maint: unflagged free DOI (link)
  18. ^ Carroll, N. REASSIGNMENT OF MONTANAZHDARCHO MINOR AS A NON-AZHDARCHID MEMBER OF THE AZHDARCHOIDEA, SVP 2015
  19. ^ Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. doi:10.1134/S0031030108060099.
  • Astibia, H.; Buffetaut, E.; Buscalioni, A.D.; Cappetta, H.; Corral, C.; Estes, R.; Garcia-Garmilla, F.; Jaeger, Mazin; Jimenez-Fuentes, J.J.; Loeuff, J. Le; Mazin, J.M.; Orue-Etxebarria, X.; Pereda-Suberbiola, J.; Powell, J.E.; Rage, J.C.; Rodriguez-Lazaro, J.; Sanz, J.L.; Tong, H.; et al. (1991). "The fossil vertebrates from Lafio (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental fauna of Europe". Terra Nova. 2 (5): 460–466. doi:10.1111/j.1365-3121.1990.tb00103.x.
  • Bennett, S. C. (2000). "Pterosaur flight: the role of actinofibrils in wing function". Historical Biology. 14 (4): 255–284. doi:10.1080/10292380009380572.
  • Nesov, L.A. (1990). "Flying reptiles of the Jurassic and Cretaceous of the USSR and the significance of their remains for the reconstruction of palaeogeographic conditions". Bulletin of Leningrad University, Series 7, Geology and Geography (in Russian). 4 (28): 3–10.
  • Nesov, L.A. (1991). "Giant flying reptiles of the family Azhdarchidae: 11. Environment, sedirnentological conditions and preservation of remains". Bulletin of Leningrad Universitv Series 7, Geology and Geography (in Russian). 3 (21): 16–24.
Retrieved from "https://en.wikipedia.org/w/index.php?title=Azhdarchidae&oldid=687625340"