Helicopsis

Helicopsis
	Drawing of apical, apertural and umbilical view of a shell of Helicopsis striata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Family:	Geomitridae
Genus:	Helicopsis; Fitzinger, 1833
Type species
	Helix striata; O. F. Müller, 1774
Diversity
	about 9 species
Synonyms
	Costatella M. Kimakowicz, 1890; Helicella (Striatella) Westerlund, 1876 (junior objective synonym); Helicopsis (Helicopsis) Fitzinger, 1833· accepted, alternate representation; Heliocopsis Fitzinger, 1833 (incorrect subsequent spelling); Martha A. J. Wagner, 1914 (junior synonym); Striatella Westerlund, 1876; Striatinella Clessin, 1876;

Helicopsis is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Helicellinae of the family Geomitridae.^[2]^[3] ^[4]

Distribution

Distribution of the genus Helicopsis range Central and Eastern Europe to Iran.^[2]

Species

The species composition of Helicopsis was largely reviewed after revisions with methods of molecular taxonomy, many of the previously recognized species were shown to be conspecific and many other to be not related with true Helicopsis.^[5]^[2] After these revisions only 9 or 10 species can be placed into Helicopsis:^[5]^[2]

Helicopsis aelleni Hausdorf, 1996 - Northern Iran^[2]
Helicopsis anflousiana (Pallary, 1913)
Helicopsis austriaca Gittenberger, 1969 - Austria^[5]
Helicopsis buslimiana (Pollonera in Kobelt, 1898) ^[6] - Libya
Helicopsis carrossei (Pallary, 1936)
Helicopsis cereoflava (Bielz, 1851) - Romania, probably a synonym of Helicopsis lunulata^[2]
Helicopsis cypriola (Westerlund, 1889)
Helicopsis dejecta (Jan, 1832) ^[7] - Azov Sea
Helicopsis filimargo (Krynicki, 1833) - Southern and Eastern Ukraine, adjacent regions of Russia, extremely variable, previously treated as several species^[2]
Helicopsis gittenbergeri Hausdorf, 1990 - Greece^[2]
Helicopsis hungarica (Soós & H. Wagner, 1935) - Central Europe and isolated locality in Russia^[5]^[2]
Helicopsis larbiana (Pallary, 1928)
Helicopsis likharevi Schileyko, 1978 - Kopet Dag in Turkmenistan
Helicopsis lunulata (Krynicki, 1833) - Romania, Moldova, Ukraine, adjacent regions of Russia and probably Poland^[2]
Helicopsis persica Hausdorf & Bössneck, 2016 - Northern Iran^[2]
† Helicopsis phrygostriata (Oppenheim, 1919)
† Helicopsis piedmontanica Harzhauser, Neubauer & Esu in Harzhauser et al., 2015
† Helicopsis praecursor (Wenz, 1927)
Helicopsis striata (O. F. Müller, 1774) - type species,^[8] distributed only in Central Europe^[2]
Helicopsis subcalcarata (Nägele, 1903)
† Helicopsis suevica Gottschick & Wenz, 1927
Helicopsis teboudensis (Pallary, 1928)
Helicopsis welschi (Pallary, 1899)
† Helicopsis wenzi Schütt, 1985

Synonyms that were recognized as species before 21st century:

Helicopsis altenai Gasull, 1974 accepted as Xerotricha vatonniana (Bourguignat, 1867) (junior synonym)
Helicopsis arenosa (Krynicki, 1836) accepted as Helicopsis filimargo arenosa (Krynicki, 1836) accepted as Helicopsis filimargo (Krynicki, 1833)
Helicopsis dejecta (Rossmässler, 1838) - synonym of Helicopsis filimargo^[2]
Helicopsis depulsa (Pintér, 1969):^[8] synonym of Xerolenta depulsa (L. Pintér, 1969) (taxon inquirendum)
Helicopsis gigaxii Haas, 1924 is a synonym of Xerocrassa ripacurcica (Bofill, 1886)^[9]
Helicopsis instabilis (Rossmässler, 1838) - synonym of Helicopsis lunulata^[2]
Helicopsis leptocolpata (Pallary,1923): synonym of Xerosecta leptocolpata (P.M. Pallary, 1923)
Helicopsis luganica Gural-Sverlova, 2010 accepted as Helicopsis filimargo (Krynicki, 1833) (junior synonym)
Helicopsis martynovi Gural-Sverlova, 2010 accepted as Helicopsis filimargo (Krynicki, 1833) (junior synonym)
Helicopsis murcica (A. Schmidt, 1854) accepted as Xerocrassa subrogata (L. Pfeiffer, 1853) (invalid; based on a preoccupied original name)
Helicopsis paulhessei (Lindholm, 1936) - synonym of Helicopsis filimargo^[2]
Helicopsis retowskii (Clessin, 1883) - synonym of Helicopsis filimargo^[2]
Helicopsis subfilimargo Gural-Sverlova, 2010 accepted as Helicopsis filimargo (Krynicki, 1833) (junior synonym)
Helicopsis tabulae Chaper, 1855 accepted as Lyrocystis perplicata (Benson, 1851) (synonym)
Helicopsis turcica (Holten, 1802):^[8] synonym of Xeroleuca turcica (Chemnitz, 1795)

Species and synonyms that were placed into Helicopsis, but are not:^[2]

Helicopsis conopsis Morelet, 1876^[10]

References

^ (in German) Fitzinger L. I. (1833). "Systematisches Verzeichniß der im Erzherzogthume Oesterreich vorkommenden Weichthiere, als Prodrom einer Fauna derselben". Beiträge zur Landeskunde Oesterreich's unter der Enns 3: 88-122. Wien. page 101.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r Balashov, Igor; Neiber, Marco; Hausdorf, Bernhard (December 2020). "Phylogeny, species delimitation and population structure of the steppe-inhabiting land snail genus Helicopsis in Eastern Europe". Zoological Journal of the Linnean Society. 193 (3): 1108–1125. doi:10.1093/zoolinnean/zlaa156.
^ Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. ISSN 1055-7903. PMID 25485783.
^ MolluscaBase eds. (2022). MolluscaBase. Helicopsis Fitzinger, 1833. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=934023 on 2022-07-31
^ ^a ^b ^c ^d Duda, Michael; Haring, Elisabeth; Bieringer, Georg; Eschner, Anita; Mrkvicka, Alexander; Mason, Katharina (November 2018). "Taxonomic reassessment of Helicopsis austriaca Gittenberger, 1969 and its relationships to H. striata (O.F. Müller, 1774) and H. hungarica (Soos & H. Wagner, 1935) (Eupulmonata: Helicoidea)". Journal of Molluscan Studies: 432–450. doi:10.1093/mollus/eyy044.
^ Field Museum of Natural History Invertebrates Helicopsis buslimiana
^ Bailey-Matthews National Shell Museum Mollusk Collection: Helicopsis dejecta
^ ^a ^b ^c "Species in genus Helicopsis" (n=13). AnimalBase, accessed 11 June 2012.
^ Martínez-Ortí, A. (2011). "Xerocrassa ripacurcica". IUCN Red List of Threatened Species. 2011: e.T156456A4947695. doi:10.2305/IUCN.UK.2011-1.RLTS.T156456A4947695.en. Retrieved 16 November 2021.
^ Seddon, M.B. (1996). "Xeroleuca conopsis". IUCN Red List of Threatened Species. 1996: e.T9806A13018324. doi:10.2305/IUCN.UK.1996.RLTS.T9806A13018324.en. Retrieved 16 November 2021.

[1] (in German) Fitzinger L. I. (1833). "Systematisches Verzeichniß der im Erzherzogthume Oesterreich vorkommenden Weichthiere, als Prodrom einer Fauna derselben". Beiträge zur Landeskunde Oesterreich's unter der Enns 3: 88-122. Wien. page 101.

[Balashov-2] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r Balashov, Igor; Neiber, Marco; Hausdorf, Bernhard (December 2020). "Phylogeny, species delimitation and population structure of the steppe-inhabiting land snail genus Helicopsis in Eastern Europe". Zoological Journal of the Linnean Society. 193 (3): 1108–1125. doi:10.1093/zoolinnean/zlaa156.

[3] Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. ISSN 1055-7903. PMID 25485783.

[WoRMS-4] MolluscaBase eds. (2022). MolluscaBase. Helicopsis Fitzinger, 1833. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=934023 on 2022-07-31

[Dudaetal-5] Duda, Michael; Haring, Elisabeth; Bieringer, Georg; Eschner, Anita; Mrkvicka, Alexander; Mason, Katharina (November 2018). "Taxonomic reassessment of Helicopsis austriaca Gittenberger, 1969 and its relationships to H. striata (O.F. Müller, 1774) and H. hungarica (Soos & H. Wagner, 1935) (Eupulmonata: Helicoidea)". Journal of Molluscan Studies: 432–450. doi:10.1093/mollus/eyy044.

[6] Field Museum of Natural History Invertebrates Helicopsis buslimiana

[7] Bailey-Matthews National Shell Museum Mollusk Collection: Helicopsis dejecta

[AB_list-8] "Species in genus Helicopsis" (n=13). AnimalBase, accessed 11 June 2012.

[9] Martínez-Ortí, A. (2011). "Xerocrassa ripacurcica". IUCN Red List of Threatened Species. 2011: e.T156456A4947695. doi:10.2305/IUCN.UK.2011-1.RLTS.T156456A4947695.en. Retrieved 16 November 2021.

[10] Seddon, M.B. (1996). "Xeroleuca conopsis". IUCN Red List of Threatened Species. 1996: e.T9806A13018324. doi:10.2305/IUCN.UK.1996.RLTS.T9806A13018324.en. Retrieved 16 November 2021.

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