Grass skippers
Grass skippers | |
---|---|
both Hesperia comma Oxfordshire | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Lepidoptera |
Family: | Hesperiidae |
Subfamily: | Hesperiinae Latreille, 1809 |
Tribes[1] | |
|
Grass skippers or banded skippers[2] are butterflies of the subfamily Hesperiinae, part of the skipper family, Hesperiidae. The subfamily was established by Pierre André Latreille in 1809.[1]
Description and distribution
With over 2,000 described species, this is the largest skipper butterfly subfamily and occurs worldwide except in New Zealand.[3] About 50 percent of grass skippers live in the Neotropics.[4] 137 species are native to North America. Around 38 species are native to Australia.[5] Genera Ochlodes and Hesperia exist exclusively in the Holarctic.[6]
They are usually orange, rust, or brown in colour and have pointed forewings.[2] Many species have dark markings or black stigmas on their forewings.[2] Most members of this subfamily have an oval antenna club with an apiculus on the tip, although Carterocephalus and Piruna do not.[7] The antennae generally has a sharp bend.
Hesperiinae larvae feed on many different types of grasses and sedges and palms, though some species are limited.[7][8]
Adults typically visit flowers and hold their wings together while feeding.[2] Hesperiinae are unique in that they hold their wings partially open while resting, with the forewings and hindwings held at different angles. This is known as the "jet-plane position".[9] Most male grass skippers perch to await females.[7]
Adults are strong fliers; they move quickly and usually in a linear direction. Some of the species, however, do flutter and these species patrol for females rather than perch.[7]
Genera incertae sedis
These[10] grass skipper genera have not yet been assigned to tribes:
- Acada
- Acerbas
- Acleros
- Actinor
- Aegiale
- Agathymus
- Alera
- Ancistroides
- Andronymus
- Ankola
- Apostictopterus fuliginosus
- Arnetta
- Artitropa
- Astictopterus
- Barca
- Caenides
- Ceratrichia
- Chondrolepis
- Creteus
- Cupitha
- Eetion
- Eogenes
- Erionota
- Fresna
- Fulda
- Galerga
- Gamia
- Gangara
- Ge geta
- Gorgyra
- Gretna
- Gyroga
- Hidari
- Hyarotis
- Hypoleucis
- Iambrix
- Idmon
- Ilma
- Isma
- Isoteinon lamprospilus
- Kedestes
- Koruthaialos
- Leona
- Lepella
- Lotongus
- Lycas
- Malaza
- Matapa
- Megathymus
- Melphina
- Meza
- Miraja
- Moltena
- Monza
- Mopala
- Notocrypta
- Oerane
- Orses
- Osmodes
- Osphantes
- Paracleros
- Parosmodes
- Pardaleodes
- Paronymus
- Pemara
- Perichares
- Perrotia
- Pirdana
- Plastingia
- Platylesches
- Ploetzia
- Prada
- Praescobura chrysomaculata
- Prosopalpus
- Pseudokerana
- Pseudopirdana
- Pseudosarbia
- Psolos
- Pteroteinon
- Pudicitia
- Pyroneura
- Pyrrhopygopsis
- Quedara
- Rhabdomantis
- Salanoemia
- Scobura
- Semalea
- Stallingsia
- Stimula
- Suada
- Suastus
- Teniorhinus
- Tiacellia
- Tsitana
- Turnerina
- Udaspes
- Unkana
- Xanthodisca
- Xanthoneura
- Zela
- Zographetus
- Zophopetes
Conservation
The following grass skippers are considered at risk.[11]
Species | Status | Location |
---|---|---|
Silver spotted skipper | Rare, Protected (UK) | Europe |
Lulworth skipper | Protected (UK) | Europe |
Orange grass-dart skipper | Rare | Australia |
White-banded grass-dart skipper | Rare | Australia |
Arogos skipper | Endangered (USA states IL, NJ, NY), Threatened (USA state MN), Species of Concern (USA state IA) | North America |
Byssus skipper | Endangered (USA Federal listing) | North America |
Carson wandering skipper | Endangered (USA Federal listing) | North America |
Dukes' skipper | Threatened (US state (MI) | North America |
Dakota skipper | Threatened (Canada), Endangered (Canada province MB), Federal Candidate (USA), Vulnerable (ICUN) | North America |
Mardon skipper | Endangered (USA state WA & Federal Candidate USA) | North America |
Otto skipper | Endangered (Canada), Threatened (Canada province MB), Threatened (USA states (IL, MI, MN) | North America |
Pawnee montane skipper | Threatened (USA Federal listing) | North America |
Poweshiek ling skipper | Threatened (Canada), Endangered (USA state WI), Threatened (USA state IA, MI), Species of Concern (USA state MN) | North America |
Rare skipper | Endangered (US state DE) | North America |
Wandering skipper | Threatened (Mexico, USA) | North America |
References
- ^ a b Brower, Andrew V.Z.; Warren, Andrew. "Hesperiinae Latreille 1809". The Tree of Life Web Project. Retrieved 5 October 2018.
- ^ a b c d Daniels, Jaret C. (2003). Butterflies of the Carolinas. Cambridge, MN: Adventure Publications, Inc. p. 9. ISBN 1-59193-007-3.
- ^ Boggs, Carol L.; Watt, Ward B.; Ehrlich, Paul R., eds. (2003). Butterflies Ecology and Evolution Taking Flight. Chicago and London: The University of Chicago Press. p. 480. ISBN 0-226-06317-8.
- ^ Kükenthal, Willy (1999). Schmidt-Rhaesa, Andreas; Kristensen, Niels P. (eds.). VOLUME 1: EVOLUTION, SYSTEMATICS, AND BIOGEOGRAPHY. Berlin and New York: Walter de Gruyter. p. 274. ISBN 9-783-11015-704-8.
- ^ Kitching, R.L.; Scheermeyer, E.; Jones, R.E.; Pierce, N.E., eds. (1999). Biology of Australian Butterflies. Collingwood, Victoria, Australia: CSIRO Publishing. p. 71. ISBN 0-6430-5027-2.
- ^ Kristensen, Niels P. (1999). Lepidoptera, moths, and butterflies. Vol. 4. Berlin: De Gruyter. p. 274. ISBN 9783110157048.
- ^ a b c d Scott, James A. (1986). The Butterflies of North America A Natural History and Field Guide. Stanford University Press. p. 424-425. ISBN 0-8047-2013-4.
- ^ Minno, Marc C. (1994). Immature stages of the skipper butterflies (Lepidoptera: hesperiidae) of the United States : biology, morphology, and descriptions (PhD). University of Florida. p. 18. Retrieved 2018-09-11.
- ^ Brock, Jim P.; Kaufman, Kenn (2003). Kaufman Focus Guides Butterflies of North America. New York: Houghton Mifflin Company. p. 302. ISBN 0618254005.
- ^ Brower, Andrew V.Z.; Warren, Andrew. "Hesperiinae incertae sedis". The Tree of Life Web Project. Retrieved 5 October 2018.
- ^ Beyer, Loni Jean (2009). Oviposition Selection by a Rare Grass Skipper, Polites mardon, in Montane Habitats: Advancing Ecological Understanding for Developing Conservation Strategies (PDF) (PhD). Washington State University Vancouver. pp. 40–41. Retrieved 2018-09-16.
External links
- Markku Savela's Lepidoptera and some other life forms: Preliminary species list. Version of 6 April 2007. Retrieved 28 May 2007.
- Reference photographs: Skippers of North America Cirrus Digital Imaging
- TOL
- RMCA Images of types.
- Flickr