Bothriuridae

Bothriuridae
Cercophonius squama
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Scorpiones
Superfamily:	Scorpionoidea
Family:	Bothriuridae Simon, 1880

The Bothriuridae are a family of scorpions, comprising 151 species in 16 genera.

The family has representatives in temperate and subtropical habitats from four continents: South America, Africa, Asia, and Australia. One genus (Cercophonius) has recently been discovered in the Himalayas.

The members of this family have a unique feature - the normally pentagonal sternum consists of two transverse bars (except Liposoma and Tehuankea) and is several times broader than long.

Genus and species

Bothriurus Peters, 1861

• Bothriurus araguayae Vellard, 1934
• Bothriurus asper Pocock, 1893
• Bothriurus bertae Abalos, 1955
• Bothriurus bocki Kraepelin, 1911
• Bothriurus bonariensis (C. L. Koch, 1842)
• Bothriurus buecherli San Martín, 1934
• Bothriurus burmeisteri Kraepelin, 1894
• Bothriurus ceii Ojanguren Affilastro, 2007
• Bothriurus cerradoensis Lourenço et al., 2004
• Bothriurus chacoensis Maury & Acosta, 1993
• Bothriurus chilensis (Molina, 1782)
• Bothriurus cordubensis Acosta, 1955
• Bothriurus coriaceus Pocock, 1893
• Bothriurus dumayi (Cekalovic, 1974)
• Bothriurus flavidus Kraepelin, 1911
• Bothriurus guarani Maury, 1984
• Bothriurus huincul Mattoni, 2007
• Bothriurus illudens Mello-Leitão, 1947
• Bothriurus inermis Maury, 1981
• Bothriurus jesuita Ojanguren Affilastro, 2003
• Bothriurus keyserlingi Pocock, 1893
• Bothriurus maculatus Kraepelin, 1911
• Bothriurus mochaensis Cekalovic, 1982
• Bothriurus moojeni Mello-Leitão, 1945
• Bothriurus nendai Ojanguren Affilastro & Garcia-Mauro, 2010
• Bothriurus noa Maury, 1984
• Bothriurus olaen Acosta, 1997
• Bothriurus pampa Ojanguren Affilastro, 2002
• Bothriurus patagonicus Maury, 1968
• Bothriurus pichicuy Mattoni, 2002
• Bothriurus picunche Mattoni, 2002
• Bothriurus pora Mattoni & Acosta, 2005
• Bothriurus prospicuus Mello-Leitão, 1932
• Bothriurus rochai Mello-Leitão, 1932
• Bothriurus rochensis San Martín, 1965
• Bothriurus rubescens Mello-Leitão, 1947
• Bothriurus sanctacrucis Mattoni, 2007
• Bothriurus signatus Pocock, 1893
• Bothriurus sooretamensis San Martín, 1966
• Bothriurus trivittatus Werner, 1939
• Bothriurus vachoni San Martín, 1968
• Bothriurus vittatus (Guerin Meneville, 1838)
• Bothriurus voyati Maury, 1973
• Bothriurus ypsilon Mello-Leitão, 1935

Brachistosternus Pocock, 1893

• Brachistosternus aconcagua Ojanguren Affilastro & Luisa-Scioscia, 2007
• Brachistosternus alienus Lönnberg, 1898
• Brachistosternus andinus Chamberlin, 1916
• Brachistosternus angustimanus Ojanguren Affilastro & Roig Alsina, 2001
• Brachistosternus artigasi Cekalovic, 1974
• Brachistosternus castroi Mello-Leitão, 1940
• Brachistosternus cekalovici Ojanguren Affilastro, 2005
• Brachistosternus cepedai Ojanguren Affilastro et al., 2007
• Brachistosternus chango Ojanguren Affilastro et al., 2007
• Brachistosternus chilensis Kraepelin, 1911
• Brachistosternus coquimbo Ojanguren Affilastro et al., 2007
• Brachistosternus donosoi Cekalovic, 1974
• Brachistosternus ehrenbergii (Gervais, 1841)
• Brachistosternus ferrugineus (Thorell, 1876)
• Brachistosternus galianoae Ojanguren Affilastro, 2002
• Brachistosternus holmbergi Carbonell, 1923 [nomen dubium]
• Brachistosternus intermedius Lönnberg, 1902
• Brachistosternus kamanchaca Ojanguren Affilastro et al., 2007
• Brachistosternus kovariki Ojanguren Affilastro, 2003
• Brachistosternus mattonii Ojanguren Affilastro, 2005
• Brachistosternus montanus Roig Alsina, 1977
• Brachistosternus multidentatus Maury, 1984
• Brachistosternus negrei Cekalovic, 1975
• Brachistosternus ninapo Ochoa, 2004
• Brachistosternus ochoai Ojanguren Affilastro, 2004
• Brachistosternus paulae Ojanguren Affilastro, 2003
• Brachistosternus pegnai Cekalovic, 1969
• Brachistosternus pentheri Mello-Leitão, 1931
• Brachistosternus perettii Ojanguren Affilastro & Mattoni, 2006
• Brachistosternus peruvianus Toledo Piza, 1974
• Brachistosternus piacentinii Ojanguren Affilastro, 2003
• Brachistosternus prendini Ojanguren Affilastro, 2003
• Brachistosternus quiscapata Ochoa & Acosta, 2002
• Brachistosternus roigalsinai Ojanguren Affilastro, 2002
• Brachistosternus sciosciae Ojanguren Affilastro, 2002
• Brachistosternus simoneae Lourenço, 2000
• Brachistosternus telteca Ojanguren Affilastro, 2000
• Brachistosternus titicaca Ochoa & Acosta, 2002
• Brachistosternus turpuq Ochoa, 2002
• Brachistosternus weyenberghi (Thorell, 1876)
• Brachistosternus zambrunoi Ojanguren Affilastro, 2002

Brandbergia Prendini, 2003

• Brandbergia haringtoni Prendini, 2003

Brazilobothriurus Lourenço & Monod, 2000

• Brazilobothriurus pantanalensis Lourenço & Monod, 2000

Centromachetes Lønnberg, 1897

• Centromachetes obscurus Mello-Leitão, 1932
• Centromachetes pocockii (Kraepelin, 1894)
• Centromachetes titschaki (Werner, 1939)

Cercophonius Peters, 1861

• Cercophonius granulosus Kraepelin, 1908
• Cercophonius kershawi Glauert, 1930
• Cercophonius michaelseni Kraepelin, 1908
• Cercophonius queenslandae Acosta, 1990
• Cercophonius squama (Gervais, 1843)
• Cercophonius sulcatus Kraepelin, 1908

Lisposoma Lawrence, 1928

• Lisposoma elegans Lawrence, 1928
• Lisposoma joseehermana Lamoral, 1979

Orobothriurus Maury, 1975

• Orobothriurus alticola (Pocock, 1899)
• Orobothriurus ampay Ochoa & Acosta, 2003
• Orobothriurus atiquipa Ochoa & Acosta, 2002
• Orobothriurus calchaqui Ochoa et al., 2011
• Orobothriurus compagnuccii Ochoa et al., 2011
• Orobothriurus curvidigitus (Kraepelin, 1911)
• Orobothriurus famatina Acosta & Ochoa, 2001
• Orobothriurus grismadoi Ojanguren Affilastro et al., 2009
• Orobothriurus huascaran Ochoa et al., 2011
• Orobothriurus paessleri (Kraepelin, 1911)
• Orobothriurus parvus Maury, 1975
• Orobothriurus quewerukana Ochoa et al., 2011
• Orobothriurus ramirezi Ochoa et al., 2011
• Orobothriurus tamarugal Ochoa et al., 2011
• Orobothriurus wawita Acosta & Ochoa, 2000

Pachakutej Ochoa, 2004

• Pachakutej crassimanus (Maury, 1975)
• Pachakutej inca (Maury, 1975)
• Pachakutej iskay (Acosta & Ochoa, 2001)
• Pachakutej juchuicha Ochoa, 2004
• Pachakutej oscari Ochoa, 2004
• Pachakutej peruvianus (Mello-Leitão, 1948)

Phoniocercus Pocock, 1893

• Phoniocercus pictus Pocock, 1893
• Phoniocercus sanmartini Cekalovic, 1968

Rumikiru Ojanguren-Affilastro et al., 2012

• Rumikiru atacama Ojanguren-Affilastro et al., 2012
• Rumikiru lourençoi (Ojanguren Affilastro, 2003)

Tehuankea Cekalovic, 1973

• Tehuankea moyanoi Cekalovic, 1973

Thestylus Simon, 1880

• Thestylus aurantiurus Yamaguti & Pinto-da-Rocha, 2003
• Thestylus glasioui Bertkau, 1880
• Thestylus signatus Mello-Leitão, 1931 [species inquirenda]

Timogenes Simon, 1880

• Timogenes dorbignyi (Guérin Méneville, 1843)
• Timogenes elegans (Mello-Leitão, 1931)
• Timogenes haplochirus Maury & Roig Alsina, 1977
• Timogenes mapuche Maury, 1975
• Timogenes sumatranus Simon, 1880

Urophonius Pocock, 1893

• Urophonius achalensis Abalos & Hominal, 1974
• Urophonius brachycentrus (Thorell, 1876)
• Urophonius eugenicus (Mello-Leitão, 1931)
• Urophonius exochus (Penther, 1913)
• Urophonius granulatus Pocock, 1898
• Urophonius iheringi Pocock, 1893
• Urophonius mahuidensis Maury, 1973
• Urophonius martinezi Ojanguren Affilastro & Cheli, 2009
• Urophonius mondacai Ojanguren Affilastro et al., 2011
• Urophonius pizarroi Ojanguren Affilastro et al., 2010
• Urophonius somuncura Acosta, 2003
• Urophonius transandinus Acosta, 1998
• Urophonius tregualemuensis Cekalovic, 1981
• Urophonius tumbensis Cekalovic, 1981

Vachonia Abalos, 1954

• Vachonia martinezi Abalos, 1954