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Handkea utriformis
Scientific classification
Kingdom: Fungi
Division: Basidiomycota
Class: Basidiomycetes
Order: Agaricales
Family: Agaricaceae
Genus: Handkea
Type species
Handkea utriformis
(Bull.) Kreisel

Handkea is a genus of puffball mushrooms in the family Agaricaceae. In 1989, German mycologist Hanns Kreisel described the genus Handkea to include species of Calvatia with distinct microscopic features, including a unique type of capillitium (coarse thick-walled hyphae in the gleba), with curvy slits instead of the usual pores.[1] Although accepted by some authors,[2] the genus concept has been rejected by others.[3][4]

Phylogenetic analyses published in 2008 shows that Handkea may be grouped in a clade along with species from several other genera, including Lycoperdon, Vascellum, Morganella, Bovistella, and Calvatia.[5]



  1. ^ Kreisel H. (1989). "Studies in the Calvatia complex (basidiomycetes)". Nova Hegwigia. 48 (3–4): 281–96. 
  2. ^ Laessøe T, Pegler DN, Spooner B (1995). British puffballs, earthstars and stinkhorns: an account of the British gasteroid fungi. Kew: Royal Botanic Gardens. ISBN 0-947643-81-8. 
  3. ^ Demoulin V, Lange M. (1990). "Calvatia turneri (Ellis et Everh.) Demoulin et M. Lange, comb. nov., the correct name for C. tatrensis Hollós". Mycotaxon 38:221–26.
  4. ^ Demoulin V. (1971). "Le genre Lycoperdon en Europe et en Amérique du Nord. Etude taxonomique et phytogéographique". PhD. thesis, Université de Liège.
  5. ^ Larsson E, Jeppson M (2008). "Phylogenetic relationships among species and genera of Lycoperdaceae based on ITS and LSU sequence data from north European taxa". Mycological Research. 112 (Pt 1): 4–22. doi:10.1016/j.mycres.2007.10.018. PMID 18207380. Retrieved 2009-04-01. 

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