Kaili Formation
The Kaili Formation is a stratigraphic formation which was deposited during the Lower and Middle Cambrian (~513 to 506 million years ago). The formation is approximately 200 meters thick and was named after the city Kaili in the Guizhou province of southwest China.
The depositional environment of the Kaili formation is not entirely known, and there are two hypotheses for its formation. It may have been a nearshore marine environment with 'normal' levels of oxygenation; or it may have been a deeper water environment further from the shore, on the open continental shelf; in this setting oxygen would not be available below the surface layers of the deposited sediment.[1] The trace fossil assemblages in the formation suggest that it was below the wave base and was reasonably well-oxygenated.[1]
Fossils
The faunal assemblage is highly diverse, comprising some 110 genera among 11 phyla; of these, some 40 genera are also found in the Burgess Shale, and some 30 are also found in the Maotianshan Shale. Trilobites and eocrinoids with hard parts that are easily preserved are the most common fossils, but many animals with only soft tissues are also preserved. For example, an arthropod similar to the Ediacaran Parvancorina of the Neoproterozoic age Ediacara Hills of South Australia has been found at the Kaili site.[2]
The middle part of the Kaili Formation, the Oryctocephalus indicus Zone, contains a Burgess Shale-type Lagerstätte with many well-preserved fossils known collectively as the Kaili Biota. In terms of age, this biota is located between the two most important and famous Cambrian Lagerstätten: the middle Lower Cambrian Maotianshan Shale (containing the Chengjiang Biota), also from China: and the Middle Cambrian Burgess Shale, known from Canada.[3][4]
Some other notable fossils discovered at Kaili are putative invertebrate eggs and embryos,[5] trace fossils of the genus Gordia (not to be confused with the Gordian worms),[1] as well as Naraoia, chancellorids, Microdictyon, Wiwaxia, and Marrella.[4]
The Kaili Formation is subdivided into three trilobite zones:
- Bathynotus holopygous–Ovatoryctocara granulata Zone
- Oryctocephalus indicus Zone
- Olenoides jialaoensis Zone[6]
Candidate for GSSP
An outcrop of the Kaili Formation, the Wuliu-Zengjiayan section, is a candidate for the GSSP for the beginning of the 5th stage of the Cambrian. The FAD of two trilobites from the formation are proposed to be the official stage boundary, Oryctocephalus indicus and Ovatoryctocara granulata. Both can be correlated with formations of similar age in Siberia and China.[7]
References
- ^ a b c Wang, Y.; Lin, J. P.; Zhao, Y. L.; Orr, P. J. (2009). "Palaeoecology of the trace fossil Gordia and its interaction with nonmineralizing taxa from the early Middle Cambrian Kaili Biota, Guizhou Province, South China". Palaeogeography Palaeoclimatology Palaeoecology. 277: 141–148. doi:10.1016/j.palaeo.2009.02.017.
- ^ Lin, J. P.; Gon, S. M.; Gehling, J. G.; Babcock, L. E.; Zhao, Y. L.; Zhang, X. L.; Hu, S. X.; Yuan, J. L.; Yu, M. Y.; Peng, J. (2006). "A Parvancorina-like arthropod from the Cambrian of South China". Historical Biology. 18 (1): 33–45. doi:10.1080/08912960500508689.
- ^ Zhang, Xingliang; Wei Liu; Yuanlong Zhao (2008). "Cambrian Burgess Shale-type Lagerstätten in South China: Distribution and significance" (PDF). Gondwana Research. 14 (1–2): 255–262. doi:10.1016/j.gr.2007.06.008.
- ^ a b ZHAO Yuanlong, YUAN Jinliang, ZHU Maoyan, YANG Ruidong, GUO Qingjun, PENG Jin, YANG Xinglian (2002). "Progress and significance in research on the early Middle Cambrian Kaili Biota, Guizhou Province,China". Progress in Natural Science. 12 (9): 649–654.
{{cite journal}}: CS1 maint: multiple names: authors list (link) - ^ Lin, J.; et al. (2006). "Silicified egg clusters from a Middle Cambrian Burgess Shale–type deposit, Guizhou, south China". Geology. 34 (12): 1037–1040. Bibcode:2006Geo....34.1037L. doi:10.1130/G23006A.1.
{{cite journal}}: Explicit use of et al. in:|author=(help)CS1 maint: extra punctuation (link) - ^ Parsley, Ronald L.; Zhao Yuanlong (2006). "Long stalked eocriuoids in basal Middle Cambrian Kaili Biota, Taijiang County, Guizhou Province, China". Journal of Paleontology. 80 (6): 1058–1071. doi:10.1666/0022-3360(2006)80[1058:LSEITB]2.0.CO;2. ISSN 0022-3360.
- ^ Rozanov, A. Yu. (2008). "The 2nd Sino-Russian Symposium on the Lower Cambrian Subdivision". Paleontological Journal. 42 (4): 441–446. doi:10.1134/S0031030108040151.
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