Leaf scars are formed naturally, often at the end of the growing season for deciduous plants when a layer of cells called the abscissa layer forms between the petiole and stem. The abscission layer acts as a point of cleavage and the leaf breaks off leaving a cleanly shaped wound that is quickly healed over with protective cork. Stipules may also leave their own scars if they are present.
Bundle scars are circular or barred regions within the leaf scar where bundles of vascular tissue that had connected the leaf and the stem broke off. The number of bundle scars in a leaf scar is sometimes used as an identifying mark as they are often consistent across a species.
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